Megacraspedus dolosellus (Zeller, 1839)

(Figs 8–9)

Ypsolophus dolosellus Zeller, 1839 . Type locality: Austria.

= Ypsolophus separatellus Fischer von Röslerstamm, 1843 . Type locality: Austria.

= Megacraspedus incertellus Rebel, 1930 . Type locality: Bulgaria.

Records. HUEMER & KARSHOLT (2018): Katun valley near Katanda village, environs of Aktash village, Kuraisky khrebet; PONOMARENKO (2019) (checklist).

Material examined. RUSSIA: ALTAI REPUBLIC: Aktash vill., 50°19′12″N, 87°36′00″E, grassy steppe, rocks, 1400 m, 11.vii.2014, 1 ♂ (Barcode NMPC-LEP-0383), J. Šumpich leg. (NMPC); the same locality but 21.vi.2015, 1 ♂, J. Šumpich leg. (NMPC); the same locality but 24.vi.2019, 8 ♂♂, J. Šumpich leg. (NMPC); Kosh-Agach Distr., Kurai env. (15 km SW), Dzhangyskol lake, 50°10′49″N; 87°44′19″E, coniferous forest/steppe, 1830 m, 24.–25.vi.2015, 1♂ (Barcode NMPC-LEP-0384), J. Šumpich leg. (NMPC); the same locality but 29.–30. vi.2019, 6 ♂♂, J. Šumpich leg. (NMPC); Shebalino Distr., Cherga vill. (8 km W), 51°34′04″N; 85°28′33″E, rocky slopes, steppe, 580 m, 4 ♂♂ (Barcode NMPC-LEP-0615), J. Šumpich leg. (NMPC).

Molecular data. A genetically extremely variable species resulted in 23 BINs (HUEMER & KARSHOLT 2018), which is a unique phenomenon within the Gelechiidae (HUEMER et al. 2020) . The mean intraspecific divergence of the barcode region is 7.5% and the maximum is 13.8% (HUEMER & KARSHOLT 2018). Altaic specimens are clustered in two branches with different BINs: BOLD:ACB3319 (n = 2; 2 public, both from Altai) [as “ BIN dolo20” in HUEMER & KARSHOLT 2018], where the average intraspecific divergence of the barcode region is 0.96% (p-dist), and BOLD:ACZ3530 (n = 8, 8 public, all from Altai) [as “ BIN dolo21” in HUEMER & KARSHOLT 2018], where the average intraspecific divergence of the barcode region is 0.03% (pdist) (maximum distance 0.16% (p-dist)). Distance between these clusters is 1.38% (p-dist).

Distribution. From Europe (not recorded in northern parts of the continent) to Central Asia (HUEMER & KARSHOLT 2018, GASTṒN & VIVES MORENO 2021). Based on the current knowledge, the Altai Mountains seem to be the easternmost area of its occurrence.