Genus Phanoperla Banks

Zwick (1982, 1986b) gave formal recognition to seven Bornean Phanoperla species and left one potentially distinct form (P. species cf maculata) without resolution. Stark & Sivec (2007) recognized two additional species ( P. flabellare, P. tuberosa) and also gave new records for P. flaveola (Klapálek) and P. pumilio (Klapálek) from East Kalimantan. The following provisional key for males will assist in recognizing the 11 nominal Bornean species.

Provisional Key to Males of Bornean Phanoperla

1 Small anterior ocellus present; largest aedeagal armature consists of a subapical ring of black hooks, incomplete on venter ……………. anomala

1’ Anterior ocellus absent; aedeagal armature variable but not as above ……………………….. 2

2 Median sensilla basiconica patch of tergum 9 well developed (Fig. 13); largest aedeagal armature consists of a pair of lateral fan‐ like groups of 3‐7 black spines (Fig. 14) ……………………………….……… 3

2’ Median sensilla basiconica patch of tergum 9 absent, or reduced to a few scattered pegs (Fig. 11); largest aedeagal armature variable but not as above (Fig. 12) ……………….……………………………….……... 5

3 Median sensilla basiconica patch of tergum 9 discretely separated from lateral patches; black scale spines on aedeagal apex cover ventrolateral surface but dorsum bare ………………….………… flabellare

3’ Median sensilla basiconica patch of tergum 9 loosely united with lateral patches (Fig. 13); black scale spines on aedeagal apex cover entire circumference (Fig. 14) …………..……………………………………….......... 4

4 Bases of hemitergal processes about as wide as basal cushions; a small spiny mound located dorsad to lateral fan clusters of large spines (Fig. 14) ………………………………………………… belalong

4’ Bases of hemitergal processes narrower than basal cushions; spiny lobes absent from area of fan clusters of large spines ……………………………….. tuberosa

5 Aedeagal armature includes a single, complete, subapical ring of large hooks …………………. bakeri

5’ Aedeagal armature with two rows or clusters of large hooks (Fig. 12) …………………………………..….. 6

6 Aedeagal armature with two close‐ set, incomplete subapical rows …………………………...…………. 7

6’ Aedeagal armature with rows or clusters widely separated (Fig. 12) ………………………………….. 8

7 Subapical rows of large hooks almost complete; a nose shaped subapical aedeagal lobe present; wing veins usually bordered with brown pigment ………..……………………………………….. nervosa

7’ Subapical rows of large hooks widely separated dorsally; nose shaped subapical lobe absent; wing veins without brown pigment borders ……………………………………………… incompleta

8 Apical aedeagal hooks form a complete ring; subapical hooks set on a pair of low membranous, dorsolateral mounds ………………….……… pumilio

8’ Apical hooks not as above ………………………….. 9

9 Apical aedeagal hooks form an incomplete row; subapical hooks form an incomplete, sometimes double ventrolateral row ……………...……. maculata

9’ Aedeagal hooks not as above ………..…………… 10

10 Ventrolateral patches of large spines asterisk‐ like (Fig. 12) …………………………………….….. astrospinata

10’ Ventrolateral patches of large spines arranged in a shortrow …………………………………….… flaveola