Tolegnaro kepleri, new species

Figures 342–370, 377

TYPES: Male holotype and female allotype from Berlese and sifted leaf litter of a tropical dry forest at an elevation of 100 m at Parc National de Namoroka, 17.8 km 329° WNW Vilanandro, 16°22′36″S, 45°19′36″E, Mahajanga, 8–12 December 2002, Fisher-Griswold Arthropod Team, deposited in CASC (PBI_OON 00036155). The species epithet is in honor of the astronomer Johannes Kepler .

DIAGNOSIS: T. kepleri can be differentiated from other Tolegnaro species by the following combination of characters: carapace side microsculpture finely reticulate (figs. 346, 361); male cheliceral paturon inner margin distal portion with long curved modified seta (figs. 348, 363); sternum with radial furrows between coxae I–II, II–III, III–IV, surface finely reticulate, microsculpture only in furrows (figs. 346, 362); leg spines absent (fig. 364).

DESCRIPTION: Male: Total length 1.8. Cephalothorax: Sternum anterior margin with interrupted transverse groove (fig. 359). Mouthparts: Paturon inner margin with recumbent, curved setae (figs. 348, 363). Abdomen: Postepigastric area setae plumose. Legs: Pale orange; tibia I unmodified. Male genital opening slightly protruded, epigastric scutum apodemes large, separated from the anterior book lung opening and not connected by a furrow. Posterior spiracles connected by a groove (fig. 352). Leg spines absent (fig. 364). Tarsi I to IV superior claw teeth not examined in detail. Trichobothria examined with SEM. Tarsal organ with two sensilla visible. Genitalia: Palp cymbium with distal patch of setae. Palp proximal segments pale orange; cymbium pale orange (figs. 347–349). Embolus dorsal lamella and ventral lamella with at least two apophyses each (figs. 367–370). Palp as in figures 365 and 366. Female (PBI_OON 0003325): Total length 1.9. As in male except as noted. Cephalothorax: Mouthparts: Cheliceral paturon inner margin with scattered setae. Legs: Tibia I Emerit’s glands present. Tarsi I to IV superior claw teeth not examined in detail. Genitalia: Genital opening slit shaped (fig. 353). Sperm sac secretory glands openings flat, extending over the receptaculum center. GAp process shaped as a twisted tube (figs. 354, 355). Genitalia as in figures 350 and 351.

MATERIAL EXAMINED: N = 69. Madagascar: Mahajanga: Parc National de Baie de Baly, 12.4 km 337° NNW Soalala, tropical dry forest, 10 m, S16.01000°, E45.26500°, Nov. 26–30, 2002, Fisher-Griswold Arthropod Team, 15 mixed (CASC PBI _ OON 3044) ; 15 mixed (CASC PBI _ OON 3357); Parc National de Namoroka, 16.9 km 317° NW Vilanandro, tropical dry forest, 100 m, S16.40666°, E45.31000°, Nov. 12–16, 2002, Fisher-Griswold Arthropod Team, 9 mixed (CASC PBI _ OON 3325) ; 2 mixed (CASC PBI _ OON 36282); Parc National de Namoroka, 17.8 km 329° WNW Vilanandro, tropical dry forest, 100 m, S16.37666°, E45.32666°, Nov. 08–12, 2002, Fisher-Griswold Arthropod Team, 32 mixed (CASC PBI _ OON 1993) . Toamasina: Forêt Clasee Andriantately, rainforest, 530 m, S18.69500°, E48.81333°, Dec. 04–07, 1998, H.J. Ratsirarson, 3 mixed (CASC PBI _ OON 3838) . Toliara: 2.7 km 302° WNW Ste. Luce, littoral forest, 20 m, S24.77166°, E47.17166°, Dec. 09–11, 1998, B.L. Fisher, 1♂ (CASC PBI _ OON 3889) ; Fort Dauphin, Mt. Vasiha, primary forest, 100 m, Feb. 07, 1995, K. Emberton, 1♀ (MRAC MT. 202. 867 PBI _ OON 8036) ; Parc National d’Andohahela, 36.7 km 341° NNW Tolagnaro, montane rainforest, 900 m, S24.76388°, E46.75166°, Jan. 21–25, 2002, Fisher-Griswold Arthropod Team, 2♀ (CASC PBI _ OON 3871) ; Reserve Integrale Naturelle d’Andohahela, parcel 1, 20.0 km SE Andranondambo, 1857 m, S24.56166°, E46.72166°, Nov. 27–Dec. 05, 1995, S. Goodman, 1♀ (FMNH PBI _ OON 36301) .