vermileo (Linnaeus, 1758)

Musca vermileo Linnaeus, 1758

Rhagio vermileo Fabricius, 1775

Nemotelus vermileo De Geer, 1776

Leptis vermileo Fabricius, 1805

Vermileo degeerii Macquart, 1834

Psammorycter vermileo Blanchard, 1840

Apogon dufouri Perris, 1852

Holotype: sex not indicated in orig. des., “southern France ” (unknown deposit).

Dist.: occurring from eastern mainland Spain to Croatia, including France, Italy, Switzerland, Germany, Austria and Slovenia (Beuk & Pape 2013; Kehlmaier 2021). SPAIN: Cabrils {41°31’N 2°22’E}; FRANCE: type was assigned to occur in “southern France”, other cited locations are Bouches-du-Rhône, La Ciotat {43°10’N 5°36’E}, Nantes [47°14’19.62’’N 1°33’17.60’’E], Les Eyzeis-de-Tayac-Sireuil {44°56’N 1°02’E}, Les Grandes Landes {45°29’N 1°01’E} and Charnay {45°53’N 4°39’E}. ITALY: the species occurs in the northern half of the mainland and Sardinia (Kehlmaier 2021); there is also a report from Giglia Island {42°21’N 10°53’E}; historic records from southern Italy including Sardinia and Sicily need to be re-evaluated; AUSTRIA: reported from Linz {48°17’N 14°18’E}; CROATIA: reported from Rovinj {45°04’N 13°40’E}, Premantura {44°47’N 13°54’E}, Lošinj {44°40’N 14°22’E}, Podaca {43°08’N 17°19’E} Dubrovnik-Neretva, Mljet {42°44’N 17°32’E} and Baška Voda {44°58’N 14°47’E}; SLOVENIA: from Osp {45°34’N 13°51’E}, Piran {45°31’N 13°34’E} and Fiesta {45°31’N 13°34’E}; SWITZERLAND: from Montorge (Valais) {46°13’N 7°20’E}; GERMANY: Weil am Rhein, gravel pit Käppelin [47°35’08.3’’N 7°37’52.6’’E]. Historic records from Bulgaria and Greece need to be re-evaluated.

Refs.: De Geer, 1752: 178–191, 261–265 (larv. biol.), Tab.V (figs.); Réaumur, 1753 (larv. biol.); Linnaeus, 1758: 590 (des.); Fabricius, 1775: 762 (redes.) as Rhagio vermile o; De Geer, 1776 (larv. biol.) as Nemotelus vermile o; Fabricius, 1781: 419–420 (as Rhagio vermileo); Fabricius, 1787: 332 (as Rhagio vermileo); Fabricius, 1805: 70 (cat.) as Leptis vermile o; Lamarck, 1816: 423 (cat.) as Rhagio vermile o; Cuvier, 1817: 618–619 (as Rhagio vermileo); Meigen, 1820: 96–97 (as Leptis vermileo); De Romand, 1833: 498–499 (larv. biol. as Leptis vermileo); Macquart 1834: 428–429 (redes.) as V. degeeri i; Blanchart, 1840: 603 (redes.), as Psammorycter vermile o; Perris, 1852: 199 (redes.) as Apogon dufour i; Rondani, 1856: 153 (cat.); Von Siebold, 1861 (larv.) biol. as Leptis vermile o; Schiner, 1862: 170–171 (as Apogon dufourii); Courtiller, 1867: 71, 73–75 (larv. biol.), 72 (figs.) as Leptis vermileo; Palm, 1869: 417 (dist.) as V. degeer i; Brauer, 1883a: 43–44 (as Vermileo degeeri); Brauer, 1883b: 114 (as Vermileo degeeri); Williston, 1886: 107 (note about spp. nomenclature); Mik, 1887: 331 (syn. discussion); Röder, 1892 (syn. discussion); Coucke, 1893: 130 (as Vermileo degeeri); Mik, 1894 (syn. discussion); Griffini, 1895: 1–2 (dist.) as Psammorcyte r; Bezzi, 1898: 28 (dist. as Psammorcyter vermileo); Kertész, 1908: 297–298 (cat.); Verral, 1909: 257 (figs.) as V. degeer i; Bequaert, 1922: 277–278 (larv. biol.); Lindner, 1925: 6 (cat., fig.); Séguy, 1926: 91, 93 (figs.), 92 (des., larv. des.); Wheeler, 1930: 144–146 (nomenclature com.), 146–149 (dist.), 150–155 (larv. biol.); Edwards, 1935: 56 (fig.); Buchner, 1940 (biol., larv., biol.); Brauns, 1954 (pupae des.); Le Faucheux, 1961 (biol.); Tsacas, 1962: 200–213 (larv. des.), 201–203, 205–207, 209–212, 214, 225, pls. II, XIII–XVI (figs.), 213–215 (larv. biol.), 216–226 (comparisons to Chrysopilus and Rhagio larv.); Hemmingsen & Nielsen, 1971: 178–181 (biol.); Chassagnard & Tsacas, 1974: 14–26 (larv. morphology, comparison to Chrysopilus auratus), 15, 18, 20, 23–24, 26 (figs.); Majer, 1988: 34 (cat.); Ebejer, 1995: 87 (wrong species report, probably V. immaculatus); Thompson & Pont, 1993: 132 (list); Ludwig et al., 1996: 10–11 (larv. des.), 11–12 (larv. biol.); Nagatomi et al., 1999: 20 (key), 23 (cat.); Ovtshinnikova, 2000: 437–438 (male gen.); Carles-Tolrá 2001: 59 (dist.); Ludwig et al., 2001: 92 (larv. des.); Carles-Tolrá & Baéz, 2002: 118 (cat.); Marchesi et al., 2005: 68 (dist.); Devetak, 2008a: 285 (dist., larv. biol.); Devetak, 2008b: 632–634 (larv. biol.); Kerr, 2010: 49,51 (figs.), 58–60, 64 (phyl.); Doczkal, 2017: 826 (dist.); Woodley & Swart, 2017: 866, 868 (figs.); Papp & Soltész, 2019: 5–7 (larv. biol.), 6 (dist.); Hubenov, 2019 (dist.), Kehlmaier, 2021: 342 (phyl., dist.).