Dasymutilla paradoxa (Gerstaecker, 1874), comb. nov.
(Figs 3, 4, 9, 10)
Mutilla paradoxa Gerstaecker 1874: 76 . Holotype, ♀, Colombia, Bogotá (ZMB, examined). Sphaerophthalma [sic!] icaris Cameron 1895: 360 . Holotype, ♀, Nicaragua, Chontales (NHM, examined), syn. nov. Sphaerophthalma [sic!] icaris: Nonveiller 1990: 87, ♀ (incertae sedis).
Mutilla paradoxa: Nonveiller 1990: 88, ♀ (incertae sedis).
Traumatomutilla icaris: Cambra 1997: 122 .
Diagnosis. FEMALE. The female of this species can be separated from other D. paradoxa species-group members by the following combination of characters: the body is predominantly black; the clypeus has a basal longitudinal tubercle; the pale setal markings of the head and mesosoma are golden; the discal spots of T2 are yellow; T2 and T3 medially clothed with interspersed golden and black setae; and the pygidium is defined by strong carinae laterally.
MALE. Unknown.
Distribution. Honduras, Nicaragua, Costa Rica, Panama, Colombia.
Material examined. Type material. Holotype of Mutilla paradoxa, ♀, COLOMBIA: Cundinamarca, Bogota, Lindig S. (ZMB) ; holotype of Sphaerophthalma [sic!] icaris, ♀, NICARAGUA, Chontales, 1903-297, Janson (NHM). Other material . COLOMBIA: Tolima, Mendez, H. Cardonol, 450m, 15.XI.1995, F. Fernandez (1♀, IaVH) . COSTA RICA: Alajuela, Bijagua, 20 km S Upala, 30.IV-6.V.1991, F.D. Parker (1♀, EMUS) ; Guanacaste: 8 km E Esparta, 1000’, 11.VII.1974, C.W. O’Brien, L. O’Brien, & G.B. Marshall (1♀, CASC) ; 3 km SE Rio Naranjo, 11-21.IV.1992, F.D. Parker (1♀, FSCA) ; same data except 1-14.IX.1993 (1♀, EMUS); same data except 24- 26.V.1993 (1♀, EMUS); Sugar Beach, 3 km N Potrero, 9-11.XI.1991, A.S. Menke (1♀, USNM) ; Heredia, Estancion Biological de La Selva, 5-8.III.2001, E.G. Riley (1♀, TAMU) . HONDURAS, San Jose, C.A., 13.IV.1923, S.C. Brumer (1♀, UMSP) . NICARAGUA, Rivas, Reserva la Flor, 25-30.I.2003, E. van den Berghe (1♀, EMUS) . PANAMA: Canal Zone, Barro Colorado Island, 4.IV.1935, W.M. Wheeler (1♀, USNM) ; same data except 7.I.1977, S. Levings (1♀, USNM); same data except 28.III.1977 (1♀, USNM); same data except 15.IV.1977 (1♀, USNM); same data except 29.III.1980, R. Silbergli (1♀, USNM); Cerro Azul, N of Tocumen, 7.VI.1958, W.J. Hanson (1♀, EMUS) .
Remarks. This species is clothed predominantly with golden setae and superficially resembles Pappognatha myrmiciformis (Cameron, 1897) and Ephuta (Ephutopsis) championi (Cameron, 1894) . These three mutillids from different genera share an overlapping distribution with the abundant ant species, Camponotus sericeiventris (Guérin-Méneville, 1838) . Yanega (1994) hypothesized that P. myrmiciforms was a Müllerian mimic of C. sericieventris; we suggest that D. paradoxa and E. championi may be as well.