Genus Indohya Beier, 1974

urn:lsid:zoobank.org:act: 41313090-4004-46AE-92DF-994574CC67AC

Indohya Beier 1974: 1003

Hyella Harvey 1993: 27–28 (synonymised by Harvey and Volschenk 2007: 386).

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Type species. Indohya: Indohya besucheti Beier, 1974, by original designation.

Hyella: Hyella humphreysi Harvey, 1993, by original designation.

Diagnosis. Species of Indohya differ from the only other genus of the family, Hya, by the absence of venom glands in the fixed chelal finger (e.g. Figs. 8C, 40C) and the juxtadentate teeth of the chelal fingers (e.g. Figs. 8C, 40C).

Remarks. The genus Indohya has a remarkable disjunct distribution in the southern hemisphere with species known from southern India, Madagascar and north-western Australia (Harvey & Volschenk 2007). This is highly suggestive of an ancient Gondwanan origin for the genus. The presence of an extensive fauna in the Pilbara—comprising epigean and troglobitic species—extends the distribution of the family into a new Australian bioregion.

The ratio sb -st / b -sb has been recently introduced in pseudoscorpion taxonomy to denote the relative position of trichobothrium sb on the movable chelal finger (Zaragoza 2017). This value was calculated for all of the new species described in this paper. Its utility, however, was found to be less useful than for chthoniid pseudoscorpions, with considerable intraspecific variation found in some species. For example, males of I. boltoni vary from 1.26 to 2.26 and females vary from 2.00 to 2.45, and males of I. lynbeazleyae vary from 1.49 to 1.89, and females vary from 0.41 to 1.87. The discrepancies within I. lynbeazleyae were evident in individuals, with significant bilateral variation. The adult specimens possess ratios as follows: male holotype (WAM T91735) with 1.49 on the left chela (with the right chela missing); male paratype (WAM T91742) with 1.71 (left) and 1.89 (right); female paratype (WAM T138709) with 0.41 (left) and 1.23 (right); female paratype (WAM T138707) with 1.10 (left) and 1.55 (right); and female paratype (WAM T138728) with 1.87 (left), but the right chela is missing from the specimen.

Key to adults of species of Indohya

1. Carapace with 18 setae arranged 4: 2: 4: 4: 4, with 4 setae in posterior row........... Indohya haroldi Harvey & Volschenk

Carapace with 12–16 setae arranged 4: 2: 4: 4: 2 or 4: 2: 4: 2: 2 or 4: 0: 4: 4: 2 or 4: 0: 4: 2: 2, with 2 setae in posterior row.. ................................................................................................... 2

2. Carapace with 16 setae arranged 4: 2: 4: 4: 2 (Fig. 55A), with 4 setae in intermediate row............................ 3

Carapace with 12–14 setae arranged 4: 2: 4: 2: 2 (e.g. Figs. 10A, 23A, 26A, 47A) or 4: 0: 4: 2: 2 (e.g. Figs. 8A, 12A, 16A, 20A, 30A, 32A, 34A), with 2 setae in intermediate row........................................................... 12

3. Carapace with 14 setae arranged 4: 0: 4: 4: 2, with 4 setae in intermediate row..................................... 4

Carapace with 12 setae arranged 4: 0: 4: 2: 2, with 2 setae in intermediate row (e.g. Figs. 8A, 16A).................... 5

4. Cheliceral hand with 5 setae; fixed chelal finger with 20–21 very flat teeth...................... Indohya besucheti Beier

Cheliceral hand with 6 setae; fixed chelal finger with 29 moderately flat teeth.................... Indohya caecata Beier

5. Larger species, e.g. chela (with pedicel) 1.90–2.01 (♂), 1.84–1.855 (♀) mm long................................... 6

Smaller species, e.g. chela (with pedicel) less than 1.0 mm long................................................ 7

6. Prolateral margin of chelal hand smooth; chelal hand (without pedicel) 1.79–1.96 (♂), 1.84 (♀) × longer than broad; setae on pedipalpal femur of moderate length, not as long as width of femur; cheliceral hand with 6 setae............................................................................................. Indohya gollum Harvey & Volschenk

Prolateral margin of chelal hand finely granulate; chelal hand (without pedicel) 1.71 (♀) × longer than broad; most setae on pedipalpal femur long, about as long as width of femur; cheliceral hand with 7 setae.................................................................................................... Indohya napierensis Harvey & Volschenk

7. Eyes present......................................................................................... 8

Eyes absent (e.g. Fig. 55A)............................................................................ 11

8. With two pairs of eyes................................................................................. 9

With one pair of eyes................................................................................. 10

9. Pedipalpal femur 0.41–0.425 mm in length; pedipalpal femur with granulations on basal half....... Indohya panops Harvey

Pedipalpal femur 0.32–0.33 mm in length; pedipalpal femur smooth........................... Indohya pusilla Harvey

10. Pedipalpal femur 0.365 (♀) mm in length; carapace with lateral setae of ocular row located on same level as median ocular setae, and median setae of intermediate row located slightly anterior to lateral setae of intermediate row................................................................................................... Indohya beieri Harvey

Pedipalpal femur 0.402 (♂), 0.467 (♀) mm in length; carapace with lateral setae of ocular row located anterior to median ocular setae, and median setae of intermediate row located anterior to lateral setae of intermediate row........................................................................................... Indohya jacquelinae Harvey & Volschenk

11. Chela (with pedicel) 3.21 (♀) × longer than broad; fixed chelal finger with 41 teeth (♀).......... Indohya typhlops Harvey

Chela (with pedicel) 3.98 (♀) × longer than broad; fixed chelal finger with 46 teeth (♀) (Fig. 55C)......................................................................................... Indohya currani Harvey & Burger, n. sp.

12. Carapace with 14 setae arranged 4: 2: 4: 2: 2, with 2 setae in ocular row (e.g. Figs. 10A, 23A, 26A, 47A, 59A, 62A)...... 13

Carapace with 12–14 setae arranged 4: 0: 4: 2: 2 or 4: 0: 4: 4: 2 (e.g. Figs. 8A, 16A), without setae in ocular row........ 18

13. Eyes present (e.g. Figs. 23A, 26B, 59A).................................................................. 14

Eyes absent (e.g. Figs. 10A, 47A, 62A)................................................................... 16

14. Basal teeth of fixed chelal finger prominent and reclined (Fig. 26F)............... Indohya cardo Harvey & Burger, n. sp.

Basal teeth of fixed chelal finger small and upright (Figs. 23C, 59D)............................................ 15

15. Larger species, e.g. chela (with pedicel) greater than 1.0 mm.................... Indohya boltoni Harvey & Burger, n. sp.

Smaller species, e.g. chela (with pedicel) less than 0.8 mm .................. Indohya julianneae Harvey & Burger, n. sp.

16. Larger species, e.g. chela (with pedicel) greater than 1.5 mm ................... Indohya adlardi Harvey & Burger, n. sp.

Smaller species, e.g. chela (with pedicel) less than 1.3 mm ................................................... 17

17. Trichobothrial ratio sb–st / sb–b = 2.47 (♀) (Fig. 62C); larger species, e.g. chela (with pedicel) 1.22–1.27 (♀) mm............................................................................. Indohya sachsei Harvey & Burger, n. sp.

Trichobothrial ratio sb–st / sb–b = 1.81 (♂) (Fig. 47C); larger species, e.g. chela (with pedicel) 1.055 (♂) mm.................................................................................... Indohya rixi Harvey & Burger, n. sp.

18. Eyes present (Fig. 60A)................................................ Indohya karenae Harvey & Burger, n. sp.

Eyes absent (e.g. Figs. 8A, 20A, 30A, 57A)............................................................... 19

19. Tergite I with 2 setae................................................................................. 20

Tergite I with 4 (or rarely 3) setae....................................................................... 22

20. Smaller species, e.g. chela (with pedicel) 1.12 mm (♂) in length................. Indohya aquila Harvey & Burger, n. sp.

Larger species, e.g. chela (with pedicel) 1.52–1.65 mm (♂), 2.09–2.18 mm (♀) in length............................ 21

21. Trichobothrium isb midway between esb and ist ...................................... Indohya humphreysi (Harvey)

Trichobothrium isb closer to esb than to ist (Fig. 8C)...................... Indohya anastomosa Harvey & Burger, n. sp.

22. Distal teeth of the fixed chelal finger rounded.............................................................. 23

At least 2 distal teeth of the fixed chelal finger pointed....................................................... 24

23. All teeth of the fixed chelal finger rounded (Fig. 20E)................... Indohya arnoldstrongi Harvey & Burger, n. sp.

At least some teeth of the fixed chelal finger pointed (Fig. 57E)................. Indohya finitima Harvey & Burger, n. sp.

24. Teeth of the fixed chelal finger rounded except for 5–7 distal teeth and occasionally the basal-most tooth (Fig. 30E).......................................................................... Indohya cockingi Harvey & Burger, n. sp.

Teeth of the fixed chelal finger including many pointed teeth (e.g. Fig. 12D, 34F)................................. 25

25. Smaller species, e.g. chela (with pedicel) less than 1.20 mm; fixed chelal finger with 46–49 teeth (Figs. 12C, 34D)....... 26

Larger species, e.g. chela (with pedicel) at least 1.39 mm; fixed chelal finger with 69–85 teeth (e.g. Figs. 32C, 42D, 49C). 27

26. Chelal hand slightly narrowed distally (Fig. 12B).......................... Indohya alexanderi Harvey & Burger, n. sp.

Chelal hand rounded (Fig. 34C)......................................... Indohya draconis Harvey & Burger, n. sp.

27. Pedipalpal femur and patella mostly granulate (Figs. 32B, 42C)................................................ 28

Pedipalpal femur and patella smooth or only partially granulate (Figs. 40B, 49B, 51B)............................. 29

28. Carapaceal setae of the median row slightly recurved (Fig. 42A)............ Indohya lynbeazleyae Harvey & Burger, n. sp.

Carapaceal setae of the median row strongly recurved (Fig. 32A)................ Indohya cribbi Harvey & Burger, n. sp.

29. Pedipalpal segments narrow, e.g. chela (with pedicel) at least 5.0 × longer than broad; femur at least 6.5 × longer than broad (Fig. 49B).......................................................................................... 30

Pedipalpal segments less narrow, e.g. chela (with pedicel) less than 4.5 × longer than broad; femur less than 6.0 × longer than broad (Figs. 40B, 51B)................................................................................ 31

30. Larger species, e.g. chela (with pedicel) 2.22–2.32 (♂), 2.22–2.55 (♀) mm......... Indohya damocles Harvey & Volschenk

Smaller species, e.g. chela (with pedicel) 1.50–1.66 (♂) mm.................. Indohya sagmata Harvey & Burger, n. sp.

31. Larger species, e.g. chela (with pedicel) 1.81 (♀) mm......................... Indohya jessicae Harvey & Burger, n. sp.

Smaller species, e.g. chela (with pedicel) 1.395 (♂), 1.505 (♀)................ Indohya scanloni Harvey & Burger, n. sp.