Tullgrenella aisenbergae Marta, sp. nov.

Figs 5B, 7M, 36A–D, 37A–D, 43D, 53A–B

Zoobank. urn:lsid:zoobank.org:act: 26A1ABE9–2F4C-491A-807A-3BA607576D6A

Type material. Holotype: ♂ (IBSP 123130), BRAZIL, São Paulo, Itirapina [22°15’26.3”S 47°49’42.3”W], 2001, R. Bertim leg. Paratypes: 1♀ (IBSP 123126), same data as holotype; 1♀ (MCN 6348), Rio Grande do Sul, Passo Fundo [28°17’11.7”S 52°25’10.7”W], 25.II.1977, A.A. Lise leg.

Other material examined. BRAZIL, São Paulo, Itirapina [22°15’26.3”S 47°49’42.3”W], 2001, R . Bertim leg., 1♀ (IBSP 123129); Rio Grande do Sul, Eldorado do Sul [30°06’10.3”S 51°39’00.7”W] , 01–11. V .1996, G. Carvalho & R. Silva leg., 1♂ (MCTP 12596); 01.XI.1996, 1♂ (MCTP 12575) .

Etymology. The name honors the Uruguayan arachnologist Anita Aisenberg, who encouraged women to continue fighting gender bias, and for her leadership in organizing the 22nd International Congress of Arachnology, Montevideo, Uruguay.

Diagnosis. Males of Tullgrenella aisenbergae Marta, sp. nov. resemble those of T. sciosciae Marta & Bustamante, sp. nov. and T. serrana by the pointed cymbium (Figs 4E vs 5C, 36C–D vs 33C–D, 38C–D), and the embolus of left palp originating at about 9’clock position, but can be distinguished from them by the pointed-down proximal retrolateral lobe of tegulum, longer conductor up to the spermophore, and the longer tegular membranous process, accompanying more than half of embolus length (Figs 36C–D). Females of T. aisenbergae Marta, sp. nov. resemble those of T. sciosciae Marta & Bustamante, sp. nov. by the C-shaped copulatory opening facing up in the medial epigynum plate (Figs 37C vs 39C), but can be distinguished from them by the longer and sclerotized copulatory ducts (Figs 7M vs 7O, 37C–D) and the posteriorly-opening coupling pocket longer than wide (Figs 7M, 37C–D).

Description. Male (IBSP 123130). Total length 5.25. Carapace length 2.50; width 1.83; height 1.00 Cephalic region length 1.25; thoracic region length 1.25. Ocular area length 1.50; Anterior eyes row 1.42 wide; Posterior eyes row 1.42 wide; Distance ALE–PME 0.18; PME–PLE 0.18. Ocular diameter AME 0.42; ALE 0.25; PME 0.06; PLE 0.15. Clypeus height 0.12. Chelicerae, paturon length 0.96; 0.48 wide; Fang 0.39 length. Endites length 0.49; 0.33 wide. Labium length 0.22; 0.29 wide. Sternum length 1.02; 0.53 wide. Abdomen length 1.92; 1.42 wide, height 0.83. Legs, femur length I: 1.66; II: 1.25; III: 1.35; IV: 1.75; patella I: 0.83; II: 0.50; III: 0.50; IV: 0.75; tibia I: 1.41; II: 0.85; III: 0.65; IV: 1.30; metatarsus I: 1.00; II: 0.75; III: 1.15; IV: 1.55; tarsus I: 0.58; II: 0.50; III: 0.55; IV: 0.7. Formula legs: 4132. Palp, length of femur 0.60, patella 0.15, tibia 0.30, cymbium 0.81. Legs macrosetae, femur I: d1–1–1, p2di; II: d1–1–1, p2di, r1; III: d1–1–1, p0–1–2, r0–1–2; IV: d1–1–1, p2di, r2di. Patella I–II: p1; III–IV: p1, r1; Tibia I: v2–2–2, p1–0–1; II: v1–2–2, p1–0–1; III: p1–1–1, r1–1–1, v0–1–2; IV: d2, v0–1–2, p1–1–1, r1–1–1 metatarsus I–II: v0–2–2; III: v0–2–2, p1–1–1, r1–1–1; IV: v0–2–2, p1–1–1di, r1–1–1di. Palp macrosetae, femur: d1–1–1. Carapace as in genus (Fig. 36A–B). Legs, femora I–II all brown, III–IV with annuli brown; patella, metatarsus, and tarsus yellow (Figs 36A–B). Abdomen as in genus. Palp, femur brown, with reduced ventral basal protuberance. Retrolateral tibial apophysis sclerotized, acute, not exceeding the ventral tegulum (Fig. 36D). Tegulum C-shaped, perpendicular to cymbium distal apex, conical pRL facing down (Fig. 36C) and with the basal portion with spermophore in one inverted U-shaped loop in the medial-ventral bulb (Fig. 36C). Embolar base projected dorsoprolaterally. Embolus thin, sclerotized, and slightly curved at the apex (9 o’clock), not exceeding the cymbium apex (Fig. 36C). Two projections on the embolus: a pars pendula, a large conductor with 2/3 of embolus length, dorsal and prolateral to the spermophore (Figs 36C–D), tegular membranous process longer and not globose (Fig. 36C, see purple arrow).

Female (MCN 6248). Total length 8.25. Carapace length 3.00; width 2.42; height 1.67. Cephalic region length 2.17; thoracic region length 0.83. Ocular area length 1.25; Anterior eyes row 1.92 wide; Posterior eyes row 1.75 wide; Distance ALE–PME 0.39; PME–PLE 0.35. Ocular diameter AME 0.50; ALE 0.33; PME 0.08; PLE 0.20. Clypeus height 0.40. Chelicerae, paturon length 1.15; 0.50 wide; Fang 0.45 length. Endites length 0.60; 0.50 wide. Labium length 0.50; 0.50 wide. Sternum length 1.25; 0.80 wide.Abdomen length 5.42; 3.08 wide, height 3.33. Legs, femur length I: 1.81; II: 1.56; III: 2.00; IV: 2.18; patella I: 1.18; II: 0.93; III: 0.93; IV: 1.00; tibia I: 1.18; II: 0.93; III: 1.06; IV: 1.75; metatarsus I: 0.93; II: 0.81; III: 1.25; IV: 1.87; tarsus I: 0.62; II: 0.62; III: 0.62; IV: 0.62. Formula legs: 4312. Palp, length of femur 0.93, patella 0.31, tibia 0.37, tarsus 0.50. Legs macrosetae, femur I–II: d1–1–1, p2di; III–IV: d1–1–1, p1di, r2di. Patella I–II: 0; III–IV: p1, r1. Tibia I: v2–2–2, p1subdi; II: v2–2; III: p2, r2, v0–1–2; IV: d2, v0–1–1, p1–1–1, r1–1–1. Metatarsus I–II: v0–2–2; III: v0–1–2, p2di, r2di, d2; IV: v0–2–2, p1–1–1di, r2di, 2d. Carapace as in genus (Figs 37A–B). Legs lighter brown and yellow. Palp yellow with dark spots on dorsum of sclerites. Abdomen: dorsum lighter brown, with yellow spots medially, and white spots proximal to posterior abdomen, venter yellow (Figs 37A–B). Spinnerets as in genus (Figs 37A–B). Epigynum: copulatory opening into a fillet, posterior-medial (Figs 5B, 7M, 37C). Copulatory ducts arranged in one monoplanar spiral, C-shaped, with one medial-dorsal loop and anteriorly arranged in the spermathecae (Figs 7M, 37C–D). Spermathecae circular, with the base of fertilization ducts placed on anterior/dorsal surface, separated by longer posteriorly-opening coupling pocket (Figs 7M, 36C). Fertilization ducts extended anteriorly to the sides (Figs 7M, 37D).

Distribution. Brazil (São Paulo and Rio Grande do Sul) (Figs 53A–B).

Natural history. Found in the Chacoan subregion (Figs 53A–B). The variation in altitudinal occurrence of this species is 97–735 m a. s. l. (Figs 53A–B).