Genus HENDELIA Czerny

Hendelia Czerny 1903: 83 . Type species, Hendelia beckeri Czerny 1903, by monotypy. Gender feminine. Becker 1905: 39 [catalogue, Palaearctic]; Kertész 1906: 320 [key]; Williston 1908: 320 [comparison to Heteroneura]; Melander & Argo 1924: 27 [family and genus revision]; Czerny 1928: 4 [family and genus revision, Palaearctic]; Schulze et al. 1930: 1517 [nomenclator]; Neave 1939b: 617 [nomenclator]; Hennig 1952: 227 [discussion of immature stages in Clusiidae]; Frey 1960: 11 (as genus and subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]; Trojan 1962: 21 [key, Poland]; Soós 1963a: 177 [redefinition, key, relationship to Prohendelia and Parahendelia], 1964a: 7 [key], 1981: 96 [Hungary, key], 1984: 11 [catalogue, Palaearctic]; Cole & Schlinger 1969: 426 [citation]; Sasakawa 1977: 238 [catalogue, Orient], 2011b: 13 [Oriental fauna]; Ferrar 1987: 122 [immature stages, illustrations of larva and puparium]; Stackelberg 1988: 505 [key, European USSR]; Greve 2005: 3 [Norway]; Sueyoshi 2006: 4 [Japan], 2014: 619 [as genus and subgenus, catalogue, Japan]; Papp et al. 2006: 190 [identified and unidentified species (including possible new subgenus), Thailand]; Lonsdale & Marshall 2007b: 120 [review, biology, redefinition, comparison to Clusiodes, synonymy of Aristohendelia Frey, Burmanochorea Frey, Labomyia Frey, Parahendelia Frey, Prohendelia Frey, Microclusaria Frey, Steleoclusiodes Frey and Xenoclusia Frey, maps for Palaearctic species], 2008b: 5 [Fiji], 2011 [revision, New world], 2010: 1044 [Manual of Central American Diptera]; Lonsdale 2009b: 230 [key to genera in Australia]; Lonsdale et al. 2010: 267 [family revision, key]; Kahanpää 2014: 288 [checklist, Finland].

Labomyia Frey 1928a: 14 . Type species, Labomyia mirabilis Frey 1928, by original designation. Gender feminine. Neave 1939b: 842 [nomenclator]; Hennig 1952: 227 [discussion of immature stages in Clusiidae]; Frey 1960: 26 [catalogue, world]; McAlpine 1960: 67 [key]; Soós 1964a: 6 [key], 1968: 4 [catalogue, Americas south of USA]; Caloren & Marshall 1998: 263 [as junior synonym of Clusiodes]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Aristohendelia Frey 1960: 14 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, World]. Gender feminine. Type species, Hendelia extensicornis Frey 1960, by original designation. Sasakawa 1977: 238 [catalogue, Orient]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Burmanochorea Frey 1960: 14 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Type species, Hendelia variabilis Frey 1960, by original designation. Gender feminine. Sasakawa 1977: 238 [catalogue, Orient]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Prohendelia Frey 1960: 13 (as subgenus) [family and genus revision, key to subgenera and species, family and genus revision, Indoaustralia, catalogue, world]. Type species, Hendelia orientalis Frey 1960, by original designation. Gender feminine. Soós 1963a: 177 [new status as genus, key, relationship to Hendelia and Parahendelia], 1964a: 7 [key]; Sasakawa 1977: 238 [catalogue, Orient]; Balthazar 1990: 417 [catalogue, Philippines]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Microclusiaria Frey 1960: 19 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Gender feminine. Type species, Clusiodes obscuripennis Frey 1960, by original designation. Sasakawa 1977: 237 [catalogue, Orient]; Balthazar 1990: 417 [catalogue, Philippines]; Papp et al. 2006: 190 [identified and unidentified species; Thailand]. Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Steleoclusiodes Frey 1960: 18 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Type species, Clusiodes nigrifrons Frey 1960, by original designation. Gender masculine. Sasakawa 1977: 237 [catalogue, Orient], 1987: 101 [key, Orient/Pacific]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Xenoclusia Frey 1960: 21 [catalogue, world]. Type species: Xenoclusia latifrons Frey 1960 [by original designation]. Gender feminine. Sasakawa 1977: 239 [catalogue, Orient]; Soós 1964a: 7 [key]; Balthazar 1990: 417 [catalogue, Philippines]; Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Parahendelia McAlpine 1960: 83 . Type species, Parahendelia latifrons McAlpine 1960, by original designation (= Hendelia armiger Lonsdale & Marshall). Gender feminine. Soós 1963a: 179 [key, relationship to Hendelia and Prohendelia], 1964a: 4 [key]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific]. Marshall 2000: 5 [as junior synonym of Clusiodes); Lonsdale & Marshall 2007b: 120 [as synonym of Hendelia].

Prophedelia. Misspelling of Prohendelia . Hua 2006: 160 [catalogue—not known from China]. Lonsdale & Marshall 2007b [citation].

aberrans (Frey) . OR: Malaysia (Sabah) , Philippines.

Clusiodes aberrans Frey 1928b: 106 . Philippines. Oriental Mindoro Province: “Calopan” [= Calapan]. HT ♀, depository unknown [originally deposited UZMH]. Delfinado & Hardy 1971: 29 [catalogue Philippine types]. Clusiodes (Microclusiaria) aberrans . Frey 1960: 19 [Philippines, catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]; Balthazar 1990: 417 [catalogue, Philippines].

Clusiodes (Clusiaria) aberrans . Sasakawa 1987: 103 [key, Orient/Pacific].

Hendelia aberrans . Lonsdale & Marshall 2007b: 147 [generic combination]; Sasakawa 2011b: 14 [Malaysia].

aenigma Frey. OR: Burma.

Hendelia (Burmanochorea) aenigma Frey 1960: 15 . Burma. NE Burma: Kambaiti, 2000m. HT ♀ UZMH. Frey 1960: 15 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].

Hendelia aenigma . Lonsdale & Marshall 2007b: 147 [list].

amerinx Lonsdale & Marshall. AU: Fiji.

Hendelia amerinx Lonsdale & Marshall 2008b: 7 . Fiji. Viti Levu: Naitasiri Province, 4km WSW Colo-i-Suva Village, Mt. Nakobalevu, 372 m. HT ♂ FNIC. Lonsdale & Marshall 2008b: 7 [illustrations of male genitalia].

angulosa (Sueyoshi) . PA: Japan.

Clusiodes angulosus Sueyoshi 2006: 6 . Japan. Honshu: Sarugajo, Kitaibaraki City, Ibaraki Prefecture, Honshu. HT ♂ BLKU. Sueyoshi 2006: 6 [map, illustrations of head and male and female genitalia, photo of wing], 2011: 7 [Japan, pictoral key], 2014: 618 [catalogue, Japan]; Lonsdale & Marshall 2007b: 154 [lapsus]. Hendelia angulosa Lonsdale & Marshall 2007b: 147 [generic combination]; Sueyoshi 2012: 27 [Japan].

arguta (McAlpine) . AU: Australia (New South Wales, Victoria).

Clusiodes arguta McAlpine 1960: 82 . Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS. Clusiodes (Clusiodes) arguta . Sasakawa 1987: 102 [key, Orient/Pacific].

Clusiodes argutus . Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific].

Hendelia arguta . Lonsdale & Marshall 2007b: 147 [generic combination].

armiger Lonsdale & Marshall. AU: Australia (New South Wales, Queensland).

Parahendelia latifrons McAlpine 1960: 84 . Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS. McAlpine 1960: 84 [illustrations of head], 1976: 75 [discussion of vibrissal structure]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific].

Clusiodes latifrons . Marshall 2000: 3 [generic combination, male agonistic behaviour, photos]. Hendelia armiger Lonsdale & Marshall 2007b: 118 [generic combination, replacement for name preoccupied by Frey (1960)].

beckeri Czerny. PA: Austraia and central Europe to Norway, Finland, eastern and northern coasts of the Black Sea, central and eastern Russia, Japan, China (Sichuan, Zeijiang)*, South Korea *.

Hendelia Beckeri Czerny 1903: 84 . Austria. Bad Hall. ST ♂♀, depository unknown. Czerny 1903: 84 [illustrations of body and heads of both sexes], 1928: 4 [redescription, illustrations of head and body, redescription of females as nigripalpis Czerny]; Becker 1905: 39 [catalogue, Palaearctic].

Hendelia beckeri . Melander & Argo 1924: 27 [summary of original description]; Tuomikoski 1936: 183 [Finland]; Trojan 1962: 21 [key, redescription, illustrations of male and female heads]; Sasakawa 1963: 18 [Japan]; Hackman 1980: 148 [checklist, Finland]; Soós 1981: 97 [Hungary, illustrations of habitus, head], 1984: 11 [catalogue, Palaearctic]; Greve 1983: 55 [habitat, Norway], 2005: 3 [considered rare, Norway]; Greve & Midtgaard 1986: 91 [Norway]; Zuska 1987: 235 [Czech Republic]; Mamaev 1987: 115 [Russian Far East]; Stackelberg 1988: 505 [key, illustration of female head, European USSR]; Ceianu 1989: 29 [catalogue, Romania]; Roháček 1995: 125 [puparia in decaying alder and birch, adults on aspen trunks, biology and habitat, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 2009: 219 [photo of male, Slovakia], 2012: 99 [central Slovakia], 2013: 128 [central Slovakia]; Merz 1998: 276 [checklist, Switzerland]; Yagi & Sawada 2000: 40 [Mamaev collection holdings]; Gärdenfors 2000: 222 [Red List of Swedish species]; Roháček et al. 2005: 280 [catalogue, habitat, considered local and rare in Czech Republic, Czech Republic]; Barták & Kubík 2005: 280 [Czech Republic]; Sueyoshi 2006: 14 [Japan], 2011: 6 [Japan, pictoral key], 2012: 28 [Japan], 2014: 619 [catalogue, Japan]; Lonsdale & Marshall 2007b: 147 [biology, illustrations of genitalia and heads, relationships]; Schacht et al. 2007: 66 [Germany]; Nilsson 2009: 5 [Sweden, illustration of female]; Adaschkiewitz 2009: 287 [Germany]; Kahanpää 2014: 288 [checklist, Finland].

Hendelia nigripalpis Czerny 1928: 5 [illustration of head]. Austria. ST ♀ [= ♀ ST of H. Beckeri Czerny], depository unknown. Frey 1928c: 108 [catalogue, Finland]; Tuomikoski 1936: 183 [as synonym of beckeri Czerny]. Hendelia (Hendelia) beckeri . Frey 1960: 13 [redescription, catalogue, World].

bisetosa (Mamaev) . PA: Russia (Far East).

Clusiodes bisetosa Mamaev 1974: 115 . Russia. Far East: Kedrovaja Pad Nature Reserve. HT ♂ MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974: 115 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].

Hendelia bisetosa . Lonsdale & Marshall 2007b: 147 [generic combination, map].

bismarckensis (Sasakawa) . AU: Papua New Guinea ( Lavongai Island).

Clusiodes bismarckensis Sasakawa 1974: 160 . Papua New Guinea. Bismarck Archipelago: Lavongai Island, Banatam. HT ♂ UZMC. Sasakawa 1974: 160 [illustrations of male genitalia], 1987: 101 [key, Orient/Pacific]; Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific].

Hendelia bismarckensis . Lonsdale & Marshall 2007b: 147 [generic combination].

boliviensis Lonsdale & Marshall. NT: Bolivia.

Hendelia boliviensis Lonsdale & Marshall 2011: 4 . Bolivia. La Paz Department: Caranavi, ca. 10km NW road to ENTEL tower, 1400 m, 15°46′35ʺS, 67°35′48ʺW. HT ♀ ANCB. Lonsdale & Marshall 2011: 4 [illustration of female genitalia].

campa Lonsdale & Marshall. NT: Mexico.

Hendelia campa Lonsdale & Marshall 2011: 4 . Mexico. Veracruz State: Xalapa. HT ♂ TAUI. Lonsdale & Marshall 2011: 4 [illustrations of male genitalia, photo].

chaetostylotis (Sasakawa) . AU: Indonesia (West Papua).

Clusiodes (Clusiodes) chaetostylotis Sasakawa 1987: 103 . Indonesia. West Papua: Kebar Valley, Vogelkop, W of Manokwari, 550 m. HT ♂ BPBM [Type No. 13580]. Sasakawa 1987: 103 [illustration of male genitalia]. Hendelia chaetostylotis . Lonsdale & Marshall 2007b: 147 [generic combination].

clara (McAlpine) . AU: Australia (Queensland).

Clusiodes clara McAlpine 1960: 83 . Australia. Queensland: Silkwood, near Innisfail. HT ♀ AMS. Clusiodes (Clusiodes) clara . Sasakawa 1987: 103 [key, Orient/Pacific].

Clusiodes clarus . Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific].

Hendelia clara . Lonsdale & Marshall 2007b: 147 [generic combination].

dasyta (Sasakawa) . AU: Solomon Islands (New Georgia Island).

Clusiodes (Columbiella) dasytus Sasakawa 1987: 113 . Solomon Islands. New Georgia Island: Munda, 1– 30m. HT ♂ BPBM [Type No. 13587]. Sasakawa 1987: 113 [illustration of male genitalia].

Hendelia dasyta . Lonsdale & Marshall 2007b: 147 [generic combination].

extensicornis Frey. OR: Burma, Thailand.

Hendelia (Aristohendelia) extensicornis Frey 1960: 14 . Burma. NE Burma, Kambaiti, 2000 m. ST 6♂ 3♀ UZMH. Frey 1960: 14 [illustrations of head, catalogue, world]; Papp et al. 2006: 191 [Thailand].

Hendelia extensicornis . Lonsdale & Marshall 2007b: 147 [relationships].

femorata (Sasakawa) . AU: Indonesia (Papua).

Clusiodes (Clusiodes) femoratus Sasakawa 1987: 105 . Indonesia. Papua Province: Ifar, east end of Cyclops Mts., 150 m.

HT ♀ BPBM [Type No. 13581].

Hendelia femorata . Lonsdale & Marshall 2007b: 147 [generic combination].

formosana (Hennig) . OR: Taiwan.

Clusiodes (Microclusiaria) formosana Hennig 1938: 128 . Taiwan. Paiwan District: northern Paroe. HT ♂ DEI. Clusiodes formosana . Hennig 1941: 146 [Taiwan]; Hua 2006: 160 [catalogue—not known from China]. Clusiodes (Microclusiaria) formosana . Frey 1960: 19 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]. Clusiodes (Clusiaria) formosana . Sasakawa 1987: 103 [key, Orient/Pacific].

Hendelia formosana . Lonsdale & Marshall 2007b: 147 [generic combination].

freyi (Soós) . OR: Taiwan.

Prohendelia (Prohendelia) freyi Soós 1963a: 179 . Taiwan. Kosempo. HT ♂ HNHM. Soós 1963a: 179 [illustrations of head]; Sasakawa 1977: 239 [catalogue, Orient].

Hendelia freyi . Lonsdale & Marshall 2007b: 147 [generic combination].

fulvifasciata Sasakawa. OR: Malaysia (Sabah).

Hendelia fulvifasciata Sasakawa 2011b: 14 . Malaysia. Sabah: 19 km N of Kalabakan, forest camp. HT ♂, BPBM [Type No. BPBM 17352]. Sasakawa 2011b: 14 [illustrations of male genitalia].

geniculata Sasakawa. OR: Malaysia (Sabah).

Hendelia geniculata Sasakawa 2011b: 16 . Malaysia. Sabah: 9.8 km SW of Tenom, forest camp. HT ♂, BPBM [Type No. BPBM 17355]. Sasakawa 2011b: 16 [illustrations of male genitalia].

gladiator (McAlpine) . AU: Australia (New South Wales, Queensland, Tasmania, Victoria). Clusiodes gladiator McAlpine 1960: 81 . Australia. New South Wales: National Park, south of Sydney. HT ♂ AMS. McAlpine 1960: 81 [illustrations of head and wing], 1976: 75 [discussion of spiral vibrissae and sexual dimorphism, illustrations of head and male postabdomen]; Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific]; Marshall 2000: 3 [male agonistic behaviour, photos].

Clusiodes (Clusiodes) gladiator . Sasakawa 1987: 102 [key, Orient/Pacific].

Hendelia gladiator . Lonsdale & Marshall 2007b: 147 [generic combination]; Lonsdale et al. 2010: 241 [genes sequenced for phylogenetic analysis].

heliconiae Lonsdale & Marshall. NT: Bolivia, Costa Rica, Ecuador.

Hendelia heliconiae Lonsdale & Marshall 2011: 6 . Ecuador. Pichincha Province: Maquipucuna Biological Research Station, river trail, 1200 m. HT ♂ QZAC. Lonsdale & Marshall 2010: 1046 [illustration of female genitalia as Hendelia sp.], 2011: 6 [illustrations of male and female genitalia, photo].

iotoides (Sasakawa) . AU: Malaysia (Sabah).

Clusiodes iotoides Sasakawa in Sasakawa & Pong 1990: 55 . Malaysia. Sabah: Sepilok Forest Reserve. HT ♂ FRC. Sasakawa & Pong 1990: 55 [illustrations of male genitalia].

Hendelia iotoides . Lonsdale & Marshall 2007b: 147 [generic combination].

kinetrolicros (Caloren & Marshall) . NT: Costa Rica, Mexico (Chiapas, Veracruz).

Clusiodes kinetrolicros Caloren & Marshall 1998: 295 . Costa Rica. Guanacaste Province : Guanacaste Conservation Area, Cacao Field Station, 900 m. HT ♂ CNC. Caloren & Marshall 1998: 295 [illustrations of head, wing and male and female genitalia].

Hendelia kinetrolicros . Lonsdale & Marshall 2007b: 147 [generic combination], 2010: 1043 [Manual of Central American Diptera], 2011: 6 [photo, reproduction of genitalic figures].

laosica Sasakawa. OR: Laos.

Hendelia laosica Sasakawa 2011b: 16 . Laos. Vientian: Gi Sion Village, de Tha Ngone. HT ♂, BPBM [Type No. BPBM 17353]. Sasakawa 2011b: 16 [illustrations of male genitalia].

lateralis (Sasakawa) . OR: Philippines.

Prohendelia lateralis Sasakawa 1974: 153 . Philippines. Palawan: Pinigisan, 600 m. HT ♂ UZMC. Sasakawa 1974: 153 [illustrations of head and male genitalia], 1977: 239 [catalogue, Orient].

Hendelia (Prohendelia) lateralis . Balthazar 1990: 417 [catalogue, Philippines].

Hendelia lateralis . Lonsdale & Marshall 2007b: 147 [genus redefinition].

latifrons (Frey) . OR: Philippines.

Xenoclusia latifrons Frey 1960: 21 . Philippines. Mindanao: Suriago. HT ♀ UZMH. Frey 1960: 21 [catalogue, world]; Delfinado & Hardy 1971: 29 [catalogue Philippine types]; Balthazar 1990: 417 [catalogue, Philippines]. Hendelia latifrons . Lonsdale & Marshall 2007b: 147 [generic combination, illustration of head].

marginalis (Sasakawa) . AU: Papua New Guinea (Bougainville Island), Indonesia (Biak Island). Clusiodes (Clusiodes) marginalis Sasakawa 1987: 106 . Papua New Guinea. Bougainville Island: Kokure, 690 m. HT ♂ BPBM [Type No. 13582]. Sasakawa 1987: 106 [illustration of male genitalia]. Hendelia marginalis . Lonsdale & Marshall 2007b: 147 [generic combination].

masneri Lonsdale & Marshall. NT: Costa Rica.

Hendelia masneri Lonsdale & Marshall 2011: 8 . Costa Rica. Puntarenas Province: Monteverde Research Station, 1550 m. HT ♂ INBio. Lonsdale & Marshall 2011: 8 [illustrations of male genitalia].

megaspilos (McAlpine) . AU: Australia (New South Wales, Queensland).

Clusiodes megaspilos McAlpine 1960: 83 . Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS.

Sasakawa 1987: 102 [key, Orient/Pacific].

Hendelia megaspilos . Lonsdale & Marshall 2007b: 147 [generic combination].

megastylotis (Sasakawa) . AU: Indonesia (West Papua) , Papua New Guinea.

Clusiodes (Clusiodes) megastylotis Sasakawa 1987: 107 . Indonesia. West Papua: Manokwari, Vogelkop, 75 m. HT ♂ BPBM [Type No. 13583]. Sasakawa 1987: 107 [illustrations of male genitalia]. Hendelia megastylotis . Lonsdale & Marshall 2007b: 147 [generic combination].

melanospilus (Sasakawa) . AU: Papua New Guinea ( Bismarck Isl., Morobe) .

Clusiodes (Clusiodes) melanospilus Sasakawa 1987: 109 . Papua New Guinea. NE New Guinea: Morobe District, Wau, 1200 m. HT ♂ BPBM [Type No. 13584]. Pitkin & Evenhuis 1989: 535 [catalogue, Australasia/Pacific]. Hendelia melanospilos . Misspelling of melanospilus . Lonsdale & Marshall 2007b: 147 [generic combination].

mirabilis (Frey) . NT: Bolivia, Costa Rica, Ecuador, Peru.

Labomyia mirabilis Frey 1928a: 16 . Peru. Eastern Andes, Roque. LT 1♂ UZMH [accidental designation of lectotype by assumption of holotype (IZCN (1999) Art. 74.6) by Caloren & Marshall (1998); 2♂ ST originally described]. Frey 1960: 26 [illustration of head, catalogue, world]

Clusiodes mirabilis . Caloren & Marshall 1998: 297 [generic combination, redescription, new record (Peru), illustrations of head, wing and male and female genitalia].

Hendelia mirabilis . Lonsdale & Marshall 2007b: 147 [generic combination, habitus photos], 2010: 1043 [Manual of Central American Diptera], 2011: 9 [description of variation, photos, illustrations of male genitalia, new records (Bolivia, Costa Rica, Ecuador)].

napo (Caloren & Marshall) . NT: Ecuador, Venezuela.

Clusiodes napo Caloren & Marshall 1998: 305 . Ecuador. Napo Province: Baeza, 2000 m. HT ♂ DEBU. Caloren & Marshall 1998: 305 [illustrations of wing and male and female genitalia].

Hendelia napo . Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 12 [reproduction of genitalic illustrations, new records (Venezuela)].

nigriceps (McAlpine) . AU: Australia (New South Wales).

Parahendelia nigriceps McAlpine 1960: 85 . Australia. New South Wales: Blue Mountains, Mt. Wilson. HT ♂ AMS. McAlpine 1960: 85 [illustration of head], 1976: 75 [discussion of spiral vibrissae]; Pitkin & Evenhuis 1989: 536 [catalogue, Australasia/Pacific].

Clusiodes nigriceps . Marshall 2000: 3 [generic combination, male agonistic behaviour, photos]. Hendelia nigriceps . Lonsdale & Marshall 2007b: 147 [generic combination].

nigrifrons (Frey) . OR: Burma.

Clusiodes (Steleoclusiodes) nigrifrons Frey 1960: 18 . Burma. NE Burma: Kambaiti, 2000 m. ST 7♂ 3♀ UZMH. Frey 1960: 18 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]; Papp et al. 2006: 190 [Thailand]. Hendelia nigrifrons . Lonsdale & Marshall 2007b: 147 [generic combination].

nigripalpus Lonsdale & Marshall. NT: Bolivia.

Hendelia nigripalpus Lonsdale & Marshall 2011: 12 . Bolivia. La Paz Department: Caranavi, ca. 10 km NW, road to ENTEL tower, 1400 m, 15°46′35ʺS, 67°35′48ʺW. HT ♂ ANCB. Lonsdale & Marshall 2011: 12 [illustrations of male genitalia].

notata (Sasakawa) . OR: Malaysia (Sarawak).

Clusiodes (Clusiodes) notatus Sasakawa 1987: 111 . Malaysia. Sarawak: Kampong, Sadong, Tapuh, 300–450 m. HT ♂ [Type No. 13585], depository unknown [originally deposited in BPBM] .

Hendelia notata . Lonsdale & Marshall 2007b: 147 [generic combination]; Sasakawa 2011b: 18 [Malaysia, illustration of male genitalia].

obscuripennis (Frey) . OR: Burma, Thailand.

Clusiodes (Microclusiaria) obscuripennis Frey 1960: 19 . Burma. Tenasserim, Sukli, 75 km E of Moulmein, 600 m. HT ♀, depository unknown [originally deposited at UZMH]. Frey 1960: 19 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient]; Papp et al. 2006: 190 [Thailand].

Clusiodes (Clusiaria) obscuripennis . Sasakawa 1987: 103 [key, Orient/Pacific].

Hendelia obscuripennis . Lonsdale & Marshall 2007b: 147 [generic combination].

orientalis (Frey) . OR: Burma.

Hendelia (Prohendelia) orientalis Frey 1960: 14 . Burma. NE Burma: Kambaiti, 2000 m. ST 1♂ 2♀ UZMH. Frey 1960: 14 [catalogue, world].

Prohendelia orientalis . Soós 1963a: 182 [key, illustrations of head]; Sasakawa 1977: 239 [catalogue, Orient]. Hendelia orientalis . Lonsdale & Marshall 2007b: 147 [generic combination].

orientalis form conjuncta Frey. OR: Burma.

Hendelia (Prohendelia) orientalis form conjuncta Frey 1960: 14. Burma. Kambaiti. ST 1♂ 2♀ UZMH.

orientalis form obscurior Frey. OR: Burma.

Hendelia (Prohendelia) orientalis form obscurior Frey 1960: 14 . Burma. Kambaiti. ST 1♀ UZMH.

palliceps Frey. OR: Burma.

Hendelia (Burmanochorea) palliceps Frey 1960: 14 . Burma. NE Burma: Kambaiti, 2000 m. ST 5♂ 2♀ UZMH. Frey 1960: 14 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].

Hendelia palliceps . Lonsdale & Marshall 2007b: 147 [list].

personata Frey. OR: Burma.

Hendelia (Burmanochorea) personata Frey 1960: 15 . Burma. NE Burma: Kambaiti, 2000 m. HT ♂ UZMH. Frey 1960: 15 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].

Hendelia personata . Lonsdale & Marshall 2007b: 147 [list].

phrenzinus (Caloren & Marshall) . NT: Costa Rica, Venezuela.

Clusiodes phrenzinus Caloren & Marshall 1998: 307 . Venezuela. Aragua State: Parque Nacional Henry Pittier, Highway Maracay-Choroni, km 19, 1330 m. HT ♂ DEBU. Caloren & Marshall 1998: 307 [illustrations of wing and male and female genitalia].

Hendelia phrenzinae . Lonsdale & Marshall 2007b: 147 [generic combination, unjustified emendation]. Hendelia phrenzinus . Lonsdale & Marshall 2011: 13 [reproduction of genitalic figures, new record (Costa Rica)].

philippinensis Sasakawa. OR: Philippines.

Hendelia philippinensis Sasakawa 2011b: 18 . Philippines. Luzon: Nueva Vizcaya, Dalton Pass, 915 m. HT ♂, BPBM [Type No. BPBM 17350]. Sasakawa 2011b: 18 [illustrations of male genitalia].

plumipes (Sasakawa) . OR: Japan (Ryukyus), Malaysia (Sabah) .

Clusiodes (Clusiodes) plumipes Sasakawa 1987: 112 . Malaysia. Sabah: Ranau, 500 m. HT ♀ BPBM [Type No. 13586]. Hendelia plumipes . Lonsdale & Marshall 2007b: 147 [generic combination]; Sasakawa 2011b: 20 [Oriental revision]; Sueyoshi 2014: 619 [catalogue, Japan].

Clusiodes usikumuri Sueyoshi 2006: 11 . Japan. Ryuku: Irimote Island, Ushikumori, Taketomi Town. HT ♀ BLKU. Sueyoshi 2006: 11 [illustrations of head and female genitalia, photo of wing]; Sasakawa 2011b: 20 [as synonym of plumipes].

Hendelia usikumuri . Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 7 [Japan, pictoral key].

plumosa (Sasakawa) . OR: Japan (Ryukyus). PA: Japan (widespread).

Clusiodes (Clusiodes) plumosus Sasakawa 1964: 14 . Japan. Honshu: Ashiu, Kyoto. HT ♀ OMNH. Sasakawa 1964: 14 [lateral illustration], 1990: 540 [description of male]; Soós 1984: 12 [catalogue, Palaearctic]; Matsumoto & Sasakawa 2006: 19 [notes holotype deposition in OKAKA].

Clusiodes plumosus . Sueyoshi 2006: 9 [illustrations of head and male genitalia, photo of wing]. Hendelia plumosa . Lonsdale & Marshall 2007b: 147 [generic combination, map]; Sueyoshi et al. 2009: 101 [log emergence trap, Japan]; Sueyoshi 2011: 8 [Japan, pictoral key], 2012: 28 [Japan], 2014: 619 [catalogue, Japan].

pterygion (Caloren & Marshall) . NT: Southern Brazil.

Clusiodes pterygion Caloren & Marshall 1998: 299 . Brazil. Rio de Janeiro : Rio de Janeiro. HT ♂ AMNH. Caloren & Marshall 1998: 299 [illustrations of male genitalia and wing].

Hendelia pterygion . Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 13 [reproduction of genitalic figures].

punctifrons (Frey) . OR: Burma.

Clusiodes (Clusiodes) punctifrons Frey 1960: 18 . Burma. NE Burma: Kambaiti, 2000 m. ST 8♂ 7♀ UZMH. Frey 1960: 18 [catalogue, world]; Sasakawa 1977: 237 [catalogue, Orient], 1987: 102 [key, Orient/Pacific]. Hendelia punctifrons . Lonsdale & Marshall 2007b: 147 [generic combination].

putealis Lonsdale & Marshall. NT: Costa Rica.

Hendelia putealis Lonsdale & Marshall 2011: 15 . Costa Rica. Cartago Province: Cerro de la Muerte La Cangreja, 1950 m. HT ♀ INBio. Lonsdale & Marshall 2011: 15 [illustration of female genitalia, photo].

quatuorsetosa (Mamaev) . PA: Russia (Far East).

Clusiodes quatuorsetosa Mamaev 1974: 114 . Russia. Far East: Ussury Krai. HT ♂ [with puparium] MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974: 114 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings].

Hendelia quatuorsetosa . Lonsdale & Marshall 2007b: 147 [generic combination, map].

salebrosa Lonsdale & Marshall. NT: Bolivia.

Hendelia salebrosa Lonsdale & Marshall 2011: 15 . Bolivia. La Paz Department: Zongo, 16°06′32ʺS, 68°04′30ʺW. HT ♂ ANCB. Lonsdale & Marshall 2011: 15 [illustrations of male and female genitalia].

saopaulo (Caloren & Marshall) . NT: Brazil (São Paulo, Rio de Janeiro).

Clusiodes saopaulo Caloren & Marshall 1998: 303 . Brazil. São Paulo : São Paulo, Jaragua. HT ♂ MNRJ. Caloren & Marshall 1998: 303 [illustrations of wing and male and female genitalia].

Hendelia saopaulo . Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 16 [reproduction of genitalic figures].

similis Lonsdale & Marshall. AU: Fiji.

Hendelia similis Lonsdale & Marshall 2008b: 8 . Fiji. Viti Levu: Nukura Forest, 60– 130m. HT ♂ BPBM [Type No. 16838]. Lonsdale & Marshall 2008b: 8 [illustrations of egg and male and female genitalia, photo].

spilota Sasakawa. OR: Malaysia (Sabah).

Hendelia spilota Sasakawa 2011b: 20 . Malaysia. Sabah: 19 km N of Kalabakan. HT ♀, BPBM [Type No. BPBM 17351]. Sasakawa 2011b: 20 [illustrations of male genitalia].

stimulator (Caloren & Marshall) . NE: Mexico (Tamaulipas).

Clusiodes stimulator Caloren & Marshall 1998: 301 . Mexico. Tamaulipas State: “ Mpla ”, Gomez Farias, Rancho de Cielo. HT ♂ TAMU. Caloren & Marshall 1998: 301 [illustrations of male genitalia and wing].

Hendelia stimulator . Lonsdale & Marshall 2007b: 147 [generic combination], 2011: 16 [reproduction of genitalic figures].

unica Mamaev. PA: Russia (Far East).

Hendelia (Burmanochorea) unica Mamaev 1974: 112 . Primorsky Krai: Kedroveja [= Kedrovaya] Pad Nature Reserve. HT ♂ MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1987: 115 [Russian Far East]. Hendelia unica . Lonsdale & Marshall 2007b: 147 [list, map].

variabilis Frey. OR: Burma.

Hendelia (Burmanochorea) variabilis Frey 1960: 15 . Burma. NE Burma: Kambaiti, 2000 m. ST 1♂ 3♀ UZMH. Frey 1960: 15 [catalogue, world]; Sasakawa 1977: 238 [catalogue, Orient].

Hendelia variabilis . Lonsdale & Marshall 2007b: 147 [list].