Isoperla roguensis Szczytko & Stewart

(Figs. 14 a-d, 20 e)

Isoperla roguensis Szczytko & Stewart 1984 .

Holotype ♂, Oregon, Curry County, mouth of Rogue River.

Isoperla karuk Baumann & Lee 2009 .

Holotype ♂, California, Humboldt County, Klamath River @ junction of Aikens Creek . Syn. Sandberg, 2011b.

Male. Aedeagus: sclerotized posterior process present; body of aedeagus with one posterior lobe, one large dorsal lobe, and one anterior lobe (Fig. 14a); one bulbous and usually bi-hemispherical shaped patch of spinulae above sclerotized process and a small pair of spinule patches placed laterally (Figs. 14 a-c); sclerotized process length approaching 0.5 mm, blade-like in lateral view (Fig. 14a), distal margin expanded and slightly membranous in posterior view (Fig. 14b), shape of distal expanded margin variable, and narrow proximal portion of sclerotized process usually inverted inside posterior aedeagal membrane. Abdominal terga 8-9, 9, 9-10: without stout spinulae or long stout setae. Posterolateral margins of at least abdominal segment 8 with scale-like setae clustered in brushes of several setae. Paraprocts: curved dorsally, length if straightened subequal to combined first and second cercal segments, tapering abruptly to tiny sharp apices (Fig. 14c). Vesicle: rounded lobe, widest at base with broadly rounded apical margin (Fig. 14d).

Isoperla roguensis shares similar characters with the species of the I. marmorata complex (Table 1). See comments under I. fulva . Three distinct spinule patches distinguishes this species. Recently, Sandberg (2011b) placed I. karuk Baumann & Lee 2009 as a junior subjective synonym.

The previously described everted anterior tubelike lobes (Szczytko & Stewart 1984, fig. 55) remained inverted after live eversion, their positions indicated by a pair of small inverted areas of membrane (openings). After clearing in KOH, the internal tubelike processes were visible with openings along anteroapical margin (Fig. 20e, ventral aspect provided clearest view).

Additional Material Examined. CALIFORNIA, Plumas Co., MF Feather River, FR 12N61X access off Hwy 70, 10.5 mi (16.9 km) SE Quincy, 39.86453°N, 120.76138°W, 18/ VI /2013, J.L. York, 1♂, 1♀ (JBSC); Jamison Creek, 1.6 mi (2.6 km) E Two Rivers exit off Hwy 70, 39.813°N, 120.6821°W, 19/ VI /2013, J.L. York, 1♀ (JBSC).