Relictanum johnsoni (Curran, 1934) comb. nov.

Map 1. Figure 45 (a–e).

Baccha johnsoni Curran, 1934 . Curran, 1934: 392. Type locality: Guyana, Bartica. Holotype female MCZ. Hull, 1949a: 159 (redescription), 252 (fig. 281, female abdomen).

Ocyptamus johnsoni . Thompson et al., 1976: 20 (catalog citation).

Baccha plutonia Hull, 1948 . Hull, 1948: 2. Type locality: Peru, Lima, Iquitos. Holotype female USNM. Hull, 1949a: 166 (redescription), 194 (fig. 17, female abdomen). n. syn.

Ocyptamus plutonia . Thompson et al., 1976: 25 (catalog citation).

Baccha smarti Curran, 1939 . Curran, 1939: 10. Type locality: Guyana, Mazaruni. Holotype female BMNH. n. syn.

Ocyptamus smarti . Thompson et al., 1976: 27 (catalog citation).

Baccha trinidadensis Curran, 1939 . Curran, 1939: 11. Type locality: Trinidad, Tabaquite. Holotype female AMNH. Hull, 1949a: 212 (fig. 98, female abdomen), 254 (fig. 284, female wing). n. syn.

Ocyptamus trinidadensis . Thompson et al., 1976: 28 (catalog citation).

Male. Male unknown.

Female. Similar to R. crassum except: Face mostly black, pale reduced ventro-laterally. Frons white pollinose maculae not connected to face pollen. Vertex narrow; ocellar triangle ~2 times its length from posterior eye margin and ~1 ocellus-width from lateral eye margin. Wing hyaline but slightly dark on cells bc, c, sc, basal ⅓ of r1, r, bm and cu p. Protibia and mesotibia with basal ⅓ to ½ pale. Metatibia with basal ¼ pale, metabasitarsomere apical ¼ pale, 4th and 5th metatarsomeres slightly darker; metacoxa and metatrochanter pile entirely white. Fourth tergite sometimes sub-quadrate, slightly longer than wide. Fifth tergite sub-quadrate, slightly wider than long. Genitalia: 7th tergite rectangular, with very short baso-lateral extensions and middle slightly convex, with pile only on apical ½ and sparse to bare medially; 7th sternite absent. Eighth tergite slightly concave on posterior margin, baso-lateral corners acute and connected to 8th sternite; 8th sternite whole, with anterior notch. Tenth tergite modified and fused to cercus forming a pair of rectangular sclerites narrowly fused basally, with long narrow baso-lateral extensions, apical ½ slightly enlarged, sparse pilose on basal ¾, densely pilose and with setulae apically.

Length. 8–10.5mm; wing 6–7mm.

Distribution. Guyana, Ecuador (Napo), Peru (Cuzco, Huanuco, Lima), Trinidad.

Material examined. BOLIVIA. La Paz, Covendo, Mulford Bio Expl 1921–22, Sep, W.m. M. Mann (1 ♀, paratype Baccha plutonia, CNC Diptera 161378). GUYANA. Bartica, Type Baccha johnsoni No. Curran [red label], Collection C.W. Johnson, Type 7662 [red label], 30 Jun 1901 (1 ♀, holotype Baccha johnsoni, MCZ). ECUADOR. Napo, Coca, Napo R., 250m, 25–30 Apr 1965, L. Pena (1 ♀, CNC Diptera 161206); ..., Jatun Sacha Biol. Res. 6 km E Misahuali, 1°4'S 70°36'W, ~ 450m, land slide in forest, Malaise, 30 Apr–7 May 2002, Buck et al. (1 ♀, DEBU 00196386). PERU. Cuzco, Quincemil, 24–31 Oct 1962, [L. E.] Pena (1 ♀, CNC Diptera 161207); Huanuco, Tingo Maria, 2200ft., 14 May 1947, J. C. Pallister (1 ♀, AMNH); [Lima], Iquitos, type [red label], Type no 51373 U.S. N.M. [red label], USNM 2052087, Mar–Apr 1931 R. C. Shannon (1 ♀, holotype Baccha plutonia, USNM). TRINIDAD. Arima Val., Simla Res. Station, 25 Jun–29 Jul 1982, J. M. Carpenter & J. S. Edgerly (1 ♀, AMNH).

Comments. Although all females examined look very similar to R. crassum, they all have a sub-quadrate 5th abdominal segment and different genitalia from R. crassum .

The information about the abdominal characters of the holotype of B. smarti is from the Diptera curator at the BMNH (Nigel Wyatt). Curran (1939) described a female but stated that the holotype is a male. The holotype is actually a female, and based on Wyatt’s observations it agrees perfectly with this redescription.

Figure 281 in Hull (1949a) stated that the drawing was a male abdomen, from the holotype. This was an error, the holotype is a female.