3.2.13 Liatongus vertagus (Fabricius, 1798)

Copris vertagus Fabricius, 1798, Ent. Syst. Suppl.: 30 (ZMUC).

Oniticellus bifurcates Erichson, 1834, Nova Acta Leop. Carol., 16(2): 235, Tf. 37, Fg. 4.

Oniticellus bifurcalis White, 1844, Ann. Mag. Nat. Hist., 14: 423(BMNH); Arrow, 1931, Fauna Brit. India, Lamell., 3: 371 (Syn.).

Oniticellus dichrous Boheman, 1858, Eugenies Resa. Zool., 1: 43 (NHRS); Arrow, 1931, Fauna Brit. India, Lamell., 3: 371 (Syn.).

Oniticellus pictisternum Fairmaire, 1891, Bull. Com. Soc. Ent. Belg., 35: 193 (MNHN); Arrow, 1931, Fauna Brit. India, Lamell., 3: 371 (Syn.).

Oniticellus scutellaris Reitter, 1892, Verh. Naturforsch. Ver. Brünn, 24: 46; Arrow, 1931, Fauna Brit. India, Lamell., 3: 371 (Syn.).

Oniticellus tonkineus Fairmaire, 1893, Bull. Soc. Ent. Belg., 37: 305 (MNHN); Arrow, 1931, Fauna Brit. India, Lamell., 3: 371 (Syn.).

Oniticellus vertagus: Arrow, 1908, Ann. Mag. Nat. Hist., 8(1): 180.

Liatongus vertagus: Arrow, 1931, Fauna Brit. India, Lamell., 3: 371.

Material examined. [AFCD] China, Hong Kong, Shing Mun, 13♂ 13♀, 2010.IV.28 , coll. unknown; Tai Mo Shan, 10♂ 8♀, 2010. V.13, coll. K.H. Cheung; Grassy Hill, 6♂ 8♀, 2010.VIII.27, coll. K.H. Cheung ; Shui Hau, 2♀, 2010.XI.18, coll. C.H. Shih ; Grassy Hill, 1♂, 2010.XII.24, coll. K.H. Cheung ; Shing Mun, 4♀, 2011.IX.1, coll. K.H. Cheung ; Shing Mun, 3♂ 3♀, 2011.XI.24, coll. K.H. Cheung ; To Kwa Peng, 2♀, 2011.XII.20, coll. K.H. Cheung ; Cheung Sheung, 3♀, 2012.IV.22, coll. M. Bai ; Ma On Shan, 1♂, 2012.IV.23, coll. M. Bai ; Sam A Tsuen, 1♂ 1♀, 2012.IV.24, coll. M. Bai ; Shing Mun, 1♀, 2013.I.3, coll. K.H. Cheung ; Tai Mo Shan, 1♂ 1♀, 2015.VII.13, coll. M.H. Leung ; Grassy Hill, 3♀, 2015.VII.15, coll. M.H. Leung ; Tong Fuk, 5♀, 2015.VIII.6, coll. K.H. Cheung ; Chek Keng, 1♀, 2015.VIII.25, coll. M.H. Leung ; Tai Mo Shan, 4♀, 2015.IX.17, coll. M.H. Leung.

[ IZAS] China, Hainan, 1♂ 2♀, 1934. V.4–9, coll. Qi He; Yinggen, 200 m, 1♀, 1960. V.5, coll. Xuezhong Zhang; Tongshen, 340 m, 1♂, 1960. III.22, coll. Zhenfu Li; Jianfengling Tianchi, 100 m, 1♂, 1981. VII.3, coll. Qiuming Guo; Tongshen, 7♂ 3♀, 1997. V.26, coll. Wenzhu Li; Wuzhishan, 800 m, 2♀, 1997. V.28–31, coll. Wenzhu Li & Peiyu Yu. Yunnan, Xishuangbanna Xiaomengyang, 850 m, 8♂ 6♀, 1957. VI.16, coll. Shuyong Wang; Lancang, 1000 m, 1♀, 1957. VI.25, coll. Shuyong Wang; Xishuangbanna Mengsong, 1600 m, 6♂ 6♀, 1958.IV.24, coll. Shuyong Wang & Chunpei Hong. Xizang, Chayu Yangyu, 1♂, 1973. VII.25, coll. Fusheng Huang. Vietnam, Ha Giang, 100 m, 3♂ 12♀, 2000. XII.9, coll. Wenzhu Li; Cao Bang, 400m, 1♂ 37♀, 2000. XII.17, coll. Wenzhu Li; Lang Son, 100 m, 3♂ 3♀, 2000. XII.20, coll. Wenzhu Li; Laboratory of Hanoi University, 20 m, 9♂ 18♀, 2000. XII.23, coll. Wenzhu Li.

[BMNH] Mong Jiong, 1♂ (Type of Oniticellus bifurcalis White, 1844); Laos, Xieng Khouang, Muong Pek, 16♂ 19♀, 1918.XII.15, coll. R. V. de Salvaza ; Laos, Xieng Khouang, Ban Sai, 2♂ 3♀, 1918.XII.16, coll. R. V. de Salvaza; Luang Prabang, Muong Hai, 12♂ 14♀, 1918.IV.2, coll. R. V. de Salvaza ; Laos, Ban Houei Soui, 11♂ 10♀, 1918.I.30, coll. R. V. de Salvaza; Luang Prabang, Ban Samang, 7♂ 9♀, 1918.XI.20, coll. R. V. de Salvaza ; Tonkin, Hoa Binh, 1♀; Haut Mekong, Nam Long, 16♂ 19♀, 1918.IV.26, coll. R. V. de Salvaza .

[ MNHN] China, Yunnan, 3♂, 1913, coll. L.N. Comby; Mou-pin, 1870, coll. A. David.

Distribution. China (Hong Kong, Fujian, Guangdong, Guizhou, Hainan, Shanghai, Taiwan, Xizang, Yunnan), India, Laos, Myanmar, Vietnam.