Phyllomya takanoi Mesnil, 1970

(Figs 41, 53, 71, 85, 99, 123, 153–154)

Phyllomya takanoi Mesnil, 1970: 119 (as Phyllomyia). Holotype male (CNC, examined). Type locality: Japan, Hokkaido, Nukabira.

Phyllomya takanoi: Herting 1984: 156 (Palearctic catalog); Herting & Dely-Draskovits 1993: 388 (Palearctic catalog); Richter 2004: 368 (tachinids of Russian Far East); Shima 2014: 837 (Japanese catalog); Shima & Hara 2016: 5 (tachinids of Saitama Prefecture).

Diagnosis. Body length: 6.4–11.1 mm. Head dichoptic; no proclinate orbital setae in male, 2 in female; arista short plumose; whitish hairs on fronto-orbital plate, parafacial, pleural region of thorax, coxae, ventroproximal portion of femora, 1st abdominal sternite and venter of syntergite 1+2; no acrostichal setae or only 1 fine presutural seta; mid tibia with 2 anterodorsal setae; hind tibia with 3 preapical dorsal setae; male fore claws and pulvilli as long as 5th tarsomere; abdominal syntergite 1+2 without median discal setae.

Redescription of terminalia. Male: Sixth tergite rather long, narrowly divided into 2 hemitergites, without hairs; surstylus narrow in lateral view, narrowed to a weakly pointed apex, with several hairs on dorsal portion; cerci in dorsal view fused basally and with median longitudinal suture obliterated along basal 1/4–1/3, rather slender, evenly narrowed to apex, in lateral view weakly constricted at basal 1/3, then weakly curved ventrally; pregonite flat; postgonite extending well beyond apex of epiphallus; dorsal sclerite of distiphallus weakly curved dorsally at apex, distal membranous portion nearly as long as dorsal sclerite. Female: Sixth tergite strongly concave middorsally on anterior margin and narrowly divided into hemitergites, with several long marginal setae; 6th sternite with short fine pale yellowish hairs mixed with black hairs on posterior portion; 7th tergite divided into long ovate hemitergites, with a few hairs on marginal portion; 7th sternite with short, fine, whitish hairs mixed with black ones on posterior portion; 8th hemitergite rather long, broadly fused with hypoproct on median portion and curved anteriorly below middle; 8th sternite longer than 6th.

Description of first instar larva. Body length 0.95–1.00 mm; cephaloskeleton rather narrow on apical 1/2, dorsal margin weakly pointed near middle in lateral view; 2nd thoracic segment with 4–5 anterodorsal rows of fine spinules; 2nd to 6th abdominal segments each with 4–5 posteroventral rows of short fine spinules, without anterodorsal spinules, 7th segment with weak posterior rows of spinules.

Type material examined. Holotype ♂: Nukabira / Hokkaido / 25.vii.1964 / S. Takano (CNC).

Additional material examined. Japan, Hokkaido: 1 ♂, 3 ♀♀, Mt. Rausu, 200–900 m, 3.viii.1967, A. Nakani- shi, T. Saigusa, M. Honda & H. Shima; 2 ♂♂ , Mt. Rausu, 900–1300 m, 4.viii.1967, A. Nakanishi; 2 ♂♂, 2 ♀♀ , Akanmachi, Ashoro-cho, 31.vii.1967, H. Shima & A. Nakanishi; 1 ♂ , Ashoro, Berabonai, 5.vii.1986, H. Shima; 8 ♀♀ , Kamitoshibetsu, 10,11,16.17. viii.1996, T. Tachi (BLKU); 2 ♂♂, 2 ♀♀ , Shibecha-cho, Touro Lake, 31.vii.1997, 1.viii.1997, 14.viii.1996, T. Tachi & M. Matsumoto (BLKU); 1 ♂, 2 ♀♀ , Gojikkoku, 15.viii.1996, T. Tachi; 1 ♂, 1 ♀ , Sapporo, Jozankei, 20.viii.1996, T. Tachi (BLKU); 1 ♀ , Sapporo, Mt. Muine, 400–1000 m, 13.viii.1977, K. Ohara; 1 ♀ , Samani-cho, Mt. Apoi, 18.viii.1996, T. Tachi; Honshu: 2 ♂♂ , Iwate Pref., Matsukawa Spa, 900–1000 m., 18,24. vii.2001, H. Shima & T. Tachi; 1 ♂ , Iwate Pref., Iwateyama, 2.viii.1975, O. Yata; 5 ♀♀ , Iwate Pref., Yamagata-Vill., Hiraniwa, 15.viii.1997, M. Sueyoshi; 1 ♂ , Akita Pref., Tazawako Town, Eboshidake, 10.viii.1996, M. Sueyoshi; 3 ♀♀ , Niigata Pref., Sasagamine, Myoko-kogen, 7.viii.1997, M. Sueyoshi; 1 ♀ , Gunma Pref., Oze, 13.vii.1975, O. Yata; 1 ♂, 1 ♀ , Gunma Pref., Ozegahara, 28–30.viii.1978, K. Kurahashi; 1 ♀ , Tochigi, Nikko, Chuzenzi, 3.ix.1917, E. Gallois (CNC); 1 ♂ , Tochigi, Nikko, Yumoto, 26.vii.1973, K. Kanmiya; 1 ♂ , Saitama, Mt. Buko, 24.vii.1978, K. Hara; 1 ♂ , Saitama, Mitsumine, 16.viii.1972, K. Hara; 4 ♂♂ , Nagano Pref., Ina, Ogurogawa Vall., 1300 m, 24.vii.1998, H. Shima & T. Tachi; 1 ♂ , Nagano Pref., Mt. Azumaya, 1300–1400 m, 29.vii.1998, T. Tachi; 1 ♂, 1 ♀ , Nagano Pref., Minamimaki Vill., Mt. Yatsugatake, 1500 m, 30.vii.1998, 6.viii.1995, T. Tachi; 1 ♂ , Nagano, Okushiga, 23.vii.1986, H. Shima; 1 ♀ , Yamanashi Pref., Mt. Kitadake, 28.viii.1980, T. Goto; 3 ♂♂, 1 ♀ , Yamanashi Pref., Hirogawara, 2.viii.1995, T. Tachi; 1 ♀ , Shizuoka Pref., Mt. Fuji, 2000 m, 28.vii.1964, S. Shi- nonaga; 1 ♀ , Gifu Pref., Kamitakara Vill., Hidarimata, 1.ix.1996, M. Sueyoshi; Shikoku: 2 ♀♀ , Ehime Pref., Saijo, Iyo-fuji, 10.viii.1998, R. Matsumoto; 1 ♂ , Kochi Pref., Higashitsuno Vill., Tengukogen, 2.vii.1996, M. Sueyoshi; Kyushu: 11 ♂♂, 5 ♀♀ , Kumamoto Pref., Mt. Hakucho, 19, 29, 30.vii.1977, 2.viii.1975, K. Ohara, H. Shima, T. Saigusa & T. Goto; 2 ♂♂ , Kumamoto Pref., Gokanosho, Momiki, 23.vii.1986, A. Tanaka & K. Kanmiya. [All KUM unless otherwise indicated]

Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu).

Remarks. Shima (1988, fig. 31) illustrated the male cerci as bearing distinct mid-dorsal suture along the dorsal surface, but this is erroneous. The cerci are fused along the basal 1/2 and the mid-dorsal longitudinal suture is obliterated there.