Enicospilus vestigator (Smith, 1858) Figure 51

Ophion vestigator Smith, 1858: 122; HT ♂ from Malaysia, OUMNH, not examined.

Eniscospilus (sic) unilineatus Cameron, 1905a: 123; HT ♂ from Sri Lanka, NHMUK, examined; synonymised by Gauld and Mitchell (1981: 471).

Henicospilus xantusi Szépligeti, 1906: 138; HT ♀ from Borneo, TM, not examined; synonymised by Gauld and Mitchell (1981: 471).

Enicospilus receptor Chiu, 1954: 40; HT ♀ from Taiwan, TARI, examined; synonymised by Townes et al. (1961: 293).

Enicospilus glabrifascies Chiu, 1954: 40; HT ♂ from Taiwan, TARI, examined; synonymised by Gauld and Mitchell (1981: 471).

Enicospilus (Polycorniata) carinatus Rao and Nikam, 1971: 103; HT ♀ from India, MUC, not examined; synonymised by Gauld and Mitchell (1981: 472).

Specimens examined.

Total of 23 specimens (19♀♀4♂♂): Brunei (2♀♀), India (11♀♀1♂), Japan (5♀♀1♂), Sri Lanka (1♂), Taiwan (1♀1♂).

Type series: HT ♂ of Eniscospilus (sic) unilineatus Cameron, 1905, Peradeniya, Ceylon, SRI LANKA, P. Cameron leg. (NHMUK, Type 3b.1237); HT ♀ of Enicospilus receptor Chiu, 1954, Koshun, TAIWAN, 27.XI.1923, R. Takahashi leg. (TARI); HT ♂ of Enicospilus glabrifascies Chiu, 1954, Kuraru, TAIWAN, 1.VIII.1931, T. Shiraki leg. (TARI).

Distribution.

Australasian, Oceanic, and Oriental regions (Yu et al. 2016); new to the Eastern Palaearctic region.

Newly recorded from Japan.

JAPAN: [ Kyûshû] Nagasaki; [ Ryûkyûs] Okinawa.

Bionomics.

Unknown.

Differential diagnosis.

This species resembles E. laqueatus, E. pseudantennatus and E. tripartitus, which all have similar sclerites, but can be distinguished from E. laqueatus by the smooth outer mandibular surface (Fig. 51B) (outer mandibular surface with a diagonal setose groove in E. laqueatus, as in Fig. 24B); from E. pseudantennatus by the strongly twisted mandible (i.e., twisted by 60-80°, as in Fig. 51B, D, but 10-20° in E. pseudantennatus) and elongate discosubmarginal cell (Fig. 51F) (rather short in E. pseudantennatus); and from E. tripartitus by the strongly twisted and smooth mandible (Fig. 51B, D) (mandible rather weakly twisted and outer surface with dense and stout setae and punctures in E. tripartitus, as in Fig. 49B, D) and elongate discosubmarginal cell (Fig. 51F) (rather short in E. tripartitus, as in Fig. 49F), as summarised in Table 6.

Remarks.

Although Japanese specimens do not vary significantly, some wing characters of the holotype of E. unilineatus are different from other specimens, as Gauld and Mitchell (1981: 472) mentioned.