Mustha spinosula (Lefebvre, 1831)

Halys spinosula Lefebvre, 1831: 21; Mustha serrata Amyot & Serville, 1843: 106 (non Fabricius, 1794).

Distribution in Iran. Alborz (Wagner 1968), East Azarbaijan (Hoberlandt 1997; Havaskary et al. 2012), Fars (Safavi 1959; Farahbakhsh 1961; Hoberlandt 1997; Modarres Awal 1997b), Golestan (Hoberlandt 1955; Safavi 1959; Farahbakhsh 1961; Modarres Awal 1997b; Heiss 2002; Linnavuori 2008), Guilan, Tehran (Linnavuori 2008), Khorasan (Linnavuori 2008; Modarres Awal 2008; Mehneh et al. 2010; Nateq Golestan & Modarres Awal 2012), Kuhgiloyeh & Boyerahmad (Hoberlandt 1997), Mazandaran (Kiritshenko 1949; Farahbakhsh 1961; Modarres Awal 1997b; Sakenin et al. 2008), Semnan (Jakovlev 1877; Oshanin 1906, 1912; Farahbakhsh 1961; Modarres Awal 1997b), West Azarbaijan (Nateq Golestan 2004; Nateq Golestan et al. 2004, 2010a, b, c, 2011), Iran (no locality cited) (Rider 2006).

General distribution. Ponto-Mediterranean from Balkanic Region to Azerbaijan, Iran and Turkmenistan, Near East (Syria, Israel) (P. Moulet, unpublished data), Sinai and probably elsewhere in Egypt (Derjanschi & Péricart 2005).

Host plants. On different forest trees (Farahbakhsh 1961; Modarres Awal 1997b), deciduous trees such as Prunus sp. ( Rosaceae), Quercus sp. ( Fagaceae), also found on Cupressus ( Cupressaceae) (Linnavuori 2008), Fraxinus syriaca ( Oleaceae), Tradescantia virginiana ( Commelinaceae), Ulmus campestris ( Ulmaceae) (Sakenin et al. 2008), Crataegus sp. ( Malaceae) (Linnavuori 2008; Nateq Golestan et al. 2010a, 2011), Prunus armeniaca ( Rosaceae), Populus sp. ( Salicaceae) (Nateq Golestan et al. 2011), Juglans regia ( Juglandaceae) (Nateq Golestan et al. 2011; Nateq Golestan & Modarres Awal 2012), Artemisia sp. ( Asteraceae) (Nateq Golestan & Modarres Awal 2012); collected on Acacia ( Fabaceae) in Iran and on Olea ( Oleaceae) in Cyprus (Derjanschi & Péricart 2005).

Comments. An unknown species, Mustha sp. was recorded by Sadaghian et al. (2004) and Nikdel et al. (2011) from East Azarbaijan; we suspect that it was M. spinosula .