Lagocheilus klobukowskii (Morlet, 1885) Figs 3A, 7A, B

Cyclophorus klobukowskii Morlet, 1885[1884]: 391, 392, pl. 12, fig. 1. Type locality: Near the Kamchay rapids, around the Kébal-Réméas cave (Kampot-Hatien road); commonly found on mountains, in forests, up to Compong-Som, and on the banks of Tap-Chéang . Fischer 1973a: 46, 47.

Lagocheilus klobukowskii: Inkhavilay et al. 2019: 19, 20, figs 9b, c, 18c.

Material examined.

Locality no. 9: CUMZ-CM044 (7 shells), CUMZ-CM045 (12 specimens in ethanol). Locality no. 10: CUMZ-CM068 (3 specimens in ethanol). Locality no. 11: CUMZ-CM079 (2 shells). Locality no. 13: CUMZ-CM128 (3 shells), CM129 (53 specimens in ethanol; Fig. 3A). Locality no. 17: CUMZ-CM137 (18 shells; Fig. 7A, B). The snails were found to live on the ground among leaf litter and decaying wood, on tree trunks and limestone wall.

Distribution.

Cambodia and Laos (Inkhavilay et al. 2019).

Remarks.

This species was described from “… grotte de Kébal-Réméas (route de Kampot à Hatien) …” . We collected topotypic specimens that tend to have a variable shell colour from yellowish (Fig. 7A, see fig. 9b in Inkhavilay et al. 2019 for the syntype) to purplish-black (Fig. 7B). This limestone associated species has a wide distribution from southern Cambodia to eastern Laos (Inkhavilay et al. 2019). The snails are commonly found in montane forest, living on decaying wood, on tree trunks and exposed limestone.

Lagocheilus klobukowskii was originally placed in the genus Cyclophorus and later was transferred to the genus Lagocheilus (see Inkhavilay et al. 2019). The distinguishing characters from the genus Cyclophorus are a conic shell, an aperture thickened (not expanded), and a thick calcareous, multispiral and plate-like operculum, whereas Cyclophorus has a turbinate shell, a thick and expanded lip, and a corneous multispiral operculum.