Key to Riethia species

Adult male ( R. paluma unknown)

1 Legs banded......................................................................................... 2

- Legs pale, unbanded................................................................................... 5

2 Gonostylus short and triangular, tapering to point (Fig. 1E). Superior volsella simple lobe without digitus (Fig. 2K)............................................................................................. stictoptera Kieffer

- Gonostylus elongate oval. Superior volsella with digitus...................................................... 3

3 Digitus of superior volsella narrow at base, dilate medially. Inferior volsella with pectinate setae (Fig. 2F) neocaledonica sp.n.

- Digitus of superior volsella broadest at base, tapering. Inferior volsella lacks pectinate setae.......................... 4

4 Digitus of superior volsella broad, tapered to tooth (Fig. 2B). Wing with dark band.................... cinctipes Freeman

- Digitus of superior volsella tapered evenly to rounded apex (Fig. 2C). Wing plain, unpigmented.......... donedwardi sp.n.

5 Some setae on inner margin of gonostylus apically plumose (Fig. 1C, D)......................................... 6

- All gonostylar setae simple (Fig. 1A, B, E, F)............................................................... 7

6 Gonostylus attenuated, apex abruptly angled (Fig. 1C)............................................. phengari sp.n.

- Gonostylus tapered conventionally (Fig. 1D).................................................. plumosa Freeman

7 Digitus of superior volsella with 1 pectinate scale (Fig. 2L)...................................... wazeylandica sp.n.

- Digitus without pectinate setae........................................................................... 8

8 Inferior volsella with only simple setae (Fig. 2G).................................................. noongar sp.n.

- Inferior volsella with several pectinate scales (Fig. 2D, E, F)................................................... 9

9 Inferior volsella distally bare, with 3 pectinate scales (Fig. 1B, 2D)................................ hamodivisa sp.n.

- Inferior volsella microtrichiose, with 5–7 pectinate scales.................................................... 10

10 Digitus of superior volsella slender from base, bearing 2 long setae; inferior volsella setose from base (Fig. 2E)..................................................................................................... kakadu sp.n.

- Digitus of superior volsella broad at base, lobe bearing 3–10 short setae (Figs 2A, K, L, M). Inferior volsella bare at base..................................................................................................... 11

11 Pseudovolsella a prominent lobe (Fig. 1F, 2M). New Zealand .................................... zeylandica Freeman

- Pseudovolsella absent (no more than inner contour of gonocoxite). Australia ..................................... 12

12 Inferior volsella with pectinate setae subapical on ridge, on same plane as superior volsella (Fig. 2J)...................................................................................................... queenslandensis sp.n.

- Inferior volsella curved, broad, lacking ridge, more ventral than superior volsella (Fig. 2A).............. azeylandica sp.n.

Pupa

1. Hook row on tergite II medially divided (Fig. 3G)........................................................... 2

- Hook row on tergite II complete.......................................................................... 3

2. Conjunctival spinule bands continuous (Fig.3G). TII and III near infilled with spinules, VI with paired posterior patches of spinules (Fig. 3G)........................................................................ hamodivisa sp.n.

- Conjunctival spinule bands interrupted. TII without spinules, III with very reduced pattern, VI posteriorly bare...................................................................................................... paluma sp.n.

3. Conjunctival spinule bands on III and IV, V bare (Fig. 3D)..................................................... 4

- Conjunctival spinule bands on III, IV & V (Fig. 3E, F)....................................................... 10

4. Conjunctival spinule bands both medially interrupted (Fig. 3H)........................................ kakadu sp.n.

- Conjunctival spinule bands continuous (Fig. 3D)............................................................ 5

5. Tergite II broadly spinulose, with anterior broad transverse spinule area extending to wide postero-median area (Fig. 3D)....................................................................................................... 6

- Tergite II with few spinules, restricted to postero-median area.................................................. 9

6. Cephalic area without warts (Fig. 3A)..................................................................... 7

- Cephalic area with distinct warts (Fig. 3B)................................................................. 8

7. Tergites mid-brown. Comb with subdominant spines (Fig. 3I)..................................... azeylandica sp.n.

- Tergites almost translucent. Comb with one dominant spine (Fig. 4J)................................. phengari sp.n.

8. TVII (Fig. 4A) and all sternites bare. Cephalic warts separated at base, tapering (Fig. 3B).................. noongar sp.n.

- TVII spinulose (Fig. 4M), sternites VI–VII spinulose in posterior 1/3, VIII less so (Fig. 4N). Cephalic warts rounded, near contiguous from base to rounded apex.................................................... queenslandensis sp.n.

9. Pedes spurii B on II (Fig. 4D). Tergite II with spinule area narrower than hook row (Fig. 4D)........... stictoptera Kieffer

- Pedes spurii B absent. Tergite II with spinule area as wide as hook row (Fig. 4C)...................... plumosa Freeman spinulose (Fig. 3E)........................................................................ cinctipes Freeman - Conjunctival spinule bands clearly separated from posterior tergal spines. Tergite VIII and anal lobe with few or no spinules (Fig. 3F)........................................................................................... 11

10. Conjunctival spinule bands near merged with posterior tergal spines. Tergite VIII with posterior spinule areas, anal lobe dense

11. Conjunctival spinule bands on III, IV & V c.10 spines deep, dense and broad (Fig. 3F)................. donedwardi sp.n.

- Conjunctival spinule bands on III and IV 3–5 spinules deep, sparse and bands narrower; on V sparser, narrower (Fig. 4L)..................................................................................................... 12

12. Conjunctival spinule band on V clearly divided medially (Fig. 4L). Exuviae golden-brown, with dark apophyses. New Caledonia .................................................................................. neocaledonica sp.n.

- Conjunctival spinule band on V sparse, continuous. Exuviae pale, with indistinct apophyses......................... 13

13. Segment V with L setae simple (Fig. 4O). New Zealand ........................................ zeylandica Freeman

- Segment V with 3 L setae all taeniate. Western Australia ........................................ wazeylandica sp.n.