Macroderes greeni Kirby, 1818

(Figs. 10, 44, 58, 91)

Onthophagus greeni Kirby, 1818: 397 . Macroderes greeni: Westwood 1847: 228; Preudhomme de Borre 1880: 9; Péringuey 1901: 298; Janssens 1939: 27; Ferreira 1969: 320; Frolov & Scholtz 2005: 388.

Type locality. Types with no locality data.

Type material examined. None. Type series is deposited in BMNH (Frolov & Scholtz 2005).

Additional material examined. 1 ♂ (TMSA): “S.Afr / SW Cape Prov / Abrahamskraal farm / 33°.14' S– 18°.09' E // 25.8.1983 / E–Y: 1976 / groundtraps / 70 days / Endrody, Penrith leg // groundtraps with faeces bait // Macroderes greeni (Kirby) / det. A. Frolov / 2003”; 3 ♀♀ (TMSA): “S.Afr / S.W Cape / Struisbaai / 34°.46' S– 20°.03' E // 28. 8. 1983 // E–Y: 1987 / groundtrap / 60 days / Endrody, Penrith leg // groundtraps with faeces bait // Macroderes greeni (Kirby) / det. A. Frolov / 2003”; 1 ♂ and 7 ♀♀ (SANC): “LANGEBAAN / C P / 12 Km / SE / Farm Geelbek / A.L.V. Davis / Macroderes sp / det. CSIRO. DBRU // Macroderes greeni (Kirby) / Frolov / det / 2003 but with different dates of collecting: 1♀: 6.VII.1979; 1 ♂ and 2 ♀♀: 13.VIII.1979; 1 ♀: 23.V.1979; 1 ♀: 30.V.1979. 1 ♀: 1–29. X. 79; 1 ♀: 26. VI. 1979; 2 ♂♂ and 4 ♀♀ (SANC): “S.Afr / SW / CP / Modderrivier farm / nr Atlantis / 33° 28' S 18 20' E / 29°. VI.1987 / ALV. Davis // ex cattle dung baited / pitfall in Strandveld // Macroderes greeni (Kirby) / Frolov / det / 2003”; 1 ♀ (SAMC): “ Macroderes greeni // Macroderes greeni (Kirby) / det. Frolov / 2003”; 1 ♂ (SAMC): “Saldanha Bay // greeni Kirby // Macroderes greeni Kirby / det. Frolov / 2003 // SAM—A 043223”; 2 ♂♂ (SAMC): “Mamre-Mamlesburg Rd / Sep. 1938 // Macroderes greeni Kirby / det. Frolov / 2003 // SAM-Col-A02321”.

Size range. Males length: 9.6–11.6 mm, width: 6.2–8.0 mm; females length: 10.3–11.4 mm, width: 6.3–7.8 mm.

Differential diagnosis. This species shares the elytral punctuation shape of M. arrowi, M. foveatus and M. cornutus (Fig. 91), however, it can differentiated by the large and rounded punctures of the pronotum (Figs. 44, 58) and from the latter two species by lacking the triangular concavity in the base of the pronotum which characterise the two species.

Habitat and distribution. This species occupies a small range which extends along the south-western coast of South Africa from Cape Agulhas in the southeast and up to Saldanha Bay in the northwest (Fig. 2). The area is situated within the Southwest Fynbos Bioregion which is considered as the core of Fynbos Biome (Mucina & Rutherford 2006). The area is mostly sandstone typified by winter rains and low to very low summer rains.