Orcula Complex

Three species are included in the Orcula Complex: H. orcula, H. pantanalensis, and H. unita . These are very small to small species (ca. 1.16–1.36 mm), which have a pronotum with or without a macula, lack a pronotal scintilla, and have small plaques (Figs. 84–85, 90). The species are grouped here on the basis of the male genitalia, which have a similar general basic shape, especially in lateral aspect, and have the basal ring with a long upper extension (Figs. 88–89, 93). The currently known species are from the Pantanal region of southern Brazil ( H. pantanalensis and H. unita), and from Brazil, slightly east of the other species ( H. orcula).