Ganaspis kimorum Buffington sp. nov.

Figs 1 C, 2, 3 E, F, 4 B, D, E

Material examined.

Holotype. Japan: Tokyo Kimura Lab, field collected G. xanthopoda ‘ suzukii type’ Collected June 2010 Received USNM July 2015; female, USNMENT 00877810. Deposited in USNM . Paratypes: Japan: Tokyo Kimura Lab, field collected G. xanthopoda ‘ suzukii type’ Collected June 2010 Received USNM July 2015; USNMENT 00877811, USNMENT 00877772, USNMENT 00877823 ; Kanto Province, Tokyo district, Hachioji, Naganuma Park 35 ° 38 ' 12 " N, 139 ° 21 ' 54 " E Ex larva of Drosophila suzukii Coll. 3. VI. 2015, Leg. N. Ris & P. Girod; USNMENT 01025537 – USNMENT 01025541 ; Tokyo, Hachioji Naganuma Park 35.6368 ° N, 139.3647 ° E, ex D. suzukii on Prunus serrulata 16 Jun 2017; Girod, collector CABI colony voucher G 1 _ Tokyo; USNMENT 01867459, USNMENT 01867458, USNMENT 01025536 . China: Yunnan Province, Kunming district, Kunming, Yunnan Agricultural University, 25 ° 07 ' 41 " N, 102 ° 44 ' 50 " E, Ex larva of Drosophila suzukii Coll. 3. VI. 2015, Leg. M. Kenis; USNMENT 01025542 – USNMENT 01025546 ; Yunnan Province, Honghe district, Shiping 23 ° 41 ' 15 " N, 102 ° 32 ' 53 " E Ex larva of Drosophila suzukii Coll. 6. VI. 2015, Leg. M. Kenis; USNMENT 01025547 – USNMENT 01025551 ; Yunnan Province, Kunming district, Kunming, Yunnan Agricultural University 25 ° 07 ' 41 " N, 102 ° 44 ' 50 " E Ex larva of Drosophila suzukii Coll. 3. VI. 2015, Leg. M. Kenis; USNMENT 01025552, USNMENT 01025553; USNMENT 01734709 ; Yunnan, Kunming Xining Temple 25.1072 ° N, 102.7167 ° E, ex D. suzukii / D. pulchrella; on Prunus sp.; 17 May 2017 Kenis, collector CABI colony voucher G 1 _ Yunnan; USNMENT 01867468, USNMENT 01867467, USNMENT 01867454 – USNMENT 01867457, USNMENT 01025528 . Yunnan Province, Honghe district, Shiping 23 ° 41 ' 15 " N, 102 ° 32 ' 53 " E Ex larva of Drosophila suzukii Coll. 6. VI. 2015, Leg. M. Kenis; USNMENT 01734706, USNMENT 01734705, USNMENT 01025530, USNMENT 01734714 . Yunnan, Kunming, Dong Da Cun, Pan Long District, 25.098602 ° N 102.835000 ° E, 2239 m, ex D. suzukii on Sambucus adnata; collected 25. VII. 2016, emerged 24. VIII. 2016 Giorgino and Guerreiri DSZ 118, DSZ 137, DSZ 188; USNMENT 01025531 – USNMENT 01025533 ; Yunnan, Dali, ex D. suzukii / Sambucus williamsii; 16 Jun 2017 Kenis, collector CABI colony voucher G 1 _ Dali; USNMENT 01867465, USNMENT 01867466. Paratypes deposited in USNM .

Diagnosis.

Separated from G. brasiliensis by the completely smooth lateral aspect of the scutellum (Fig. 3 E). Separated from G. lupini by the less-developed ovipositor clip that does not extend past the midwidth of the ovipositor valve (Fig. 4 B, E). In G. lupini the ovipositor clip expands across more than half the width of the ovipositor valve (Fig. 4 C).

Description.

As in description for G. brasiliensis species complex, but with the lateral aspect of scutellum completely smooth; ovipositor clip reduced, not extending beyond the halfway point across the fused ovipositor valve. Previous studies referencing ‘ Gb G 1 ’ or ‘ G 1 ’ refer to this species (Nomano et al. 2017; Giorgini et al. 2018; Abram et al. 2022).

Etymology.

Named in honor of Prof. Kimura (Hokkaido University, retired) and Dr Kim Hoelmer (USDA-ARS, retired). The name is a combination of Kimura and Kim Hoelmer.

Cytochrome c oxidase subunit I (COI) Barcode region.

> Ganaspis _ kimorum _ BBP 857

NATATTATATTTTATTTTTGGTATTTGATCAGGAATAATTGGATCAAGTTTAAGAATAATTATTCGATTA GAATTAGGAACCCCTTCACAATTAATTAATAATGATCAAATTTATAATACAATTGTTACTACTCATGC ATTTGTAATAATTTTTTTTATAGTTATACCAATTATAGTTGGAGGATTTGGAAATTACTTAATTCCTT TAATATTATCTGCTCCTGATATATCATTCCCTCGTCTTAATAATATAAGATTTTGATTATTAATCCCT TCTTTAATTTTAATAATTTCAAGTATATTTATTGATGAAGGGTCTGGAACTGGATGAACAGTTTATCC TCCTTTATCACTAAATAAGTCCCACCCAGGAATCTCAACTGACTTAGTAATTTTTTCTCTTCATCTTA GAGGAATTTCTTCAATTTTAGGATCAATTAATTTTATTACAACTATTCTAAATATACGACCAAATTTA ATAAGTATAGATAAAATTTCTTTATTTACTTGATCCATTTTTCTTACCACTATTTTATTATTATTATC TTTACCAGTATTAGCAGGTGGAATCACTATATTACTTTTTGACCGAAATATTAATACATCTTTTTATG ACCCAATAGGAGGAGGAGACCCAATTCTATACCAACACTTATTT

> Ganaspis _ kimorum _ 233137974 _ E 02 (partial sequence, 592 bp)

ATTAGAATTAGGAACCCCTTCACAATTAATTAATAATGATCAAATTTATAATACAATTGTTACTACTCAT GCATTTGTAATAATTTTTTTTATAGTTATACCAATTATAGTTGGAGGATTTGGAAATTACTTAATTCC TTTAATATTATCTGCTCCTGATATATCATTCCCTCGTCTTAATAATATAAGATTTTGATTATTAATCC CTTCTTTAATTTTAATAATTTCAAGTATATTTATTGATGAAGGGTCTGGAACTGGATGAACAGTTTAT CCTCCTTTATCACTAAATAAGTCCCACCCAGGAATCTCAACTGACTTAGTAATTTTTTCTCTTCATCT TAGAGGAATTTCTTCAATTTTAGGATCAATTAATTTTATTACAACTATTCTAAATATACGACCAAATT TAATAAGTATAGATAAAATTTCTTTATTTACTTGATCCATTTTTCTTACCACTATTTTATTATTATTA TCTTTACCAGTATTAGCAGGTGGAATCACTATATTACTTTTTGACCGAAATATTAATACATCTTTTTA TGACCCAATAGGAGGAGGAGACCCAATTCTATACCAACACTTATTT

> Ganaspis _ kimorum _ 233137957 _ F 06

NATATTATATTTTATTTTTGGTATTTGATCAGGAATAATTGGATCAAGTTTAAGAATAATTATTCGATTA GAATTAGGAACCCCTTCACAATTAATTAATAATGATCAAATTTATAATACAATTGTTACTACTCATGC ATTTGTAATAATTTTTTTTATAGTTATACCAATTATAGTTGGAGGATTTGGACATTACTTAATTCCTT TAATATTATCTGCTCCTGATATATCATTCCCTCGTCTTAATAATATAAGATTTTGATTATTAATCCCT TCTTTAATTTTAACAATTTCAAGTATATTTATTGATGAAGGATCTGGAACCGGATGAACAGTTTATCC TCCTTTATCACTAAATAAGTCCCACCCAGGAATCTCAACTGACTTAGTAATTTTTTCTCTTCATCTTA GAGGAATTTCTTCAATTTTAGGATCAATTAATTTTATTACAACTATTCTAAATATACGACCAAATTTA ATAAGTATAGATAAAATTTCTTTATTTACTTGATCCATTTTTCTTACCACTATTTTATTATTATTATC TTTACCAGTATTAGCAGGTGGAATCACTATATTACTTTTTGACCGAAATATTAATACATCTTTTTATG ACCCAATAGGAGGAGGAGACCCAATTCTATACCAACACTTATTT

Biology.

Koinobiont endoparasitoid of Drosophila suzukii (Nomano et al. 2017; Girod et al. 2018 b; Wang et al. 2018; Seehausen et al. 2020; Daane et al. 2021). Seehausen et al. (2020) found ‘ Ganaspis cf. brasiliensis ’ G 1 could successfully parasitize D. melanogaster in lab culture.