Caecum sepimentum de Folin, 1868

Figs 15, 25D

Caecum sepimentum de Folin, 1868: 84, pl. 6, fig. 7.

Caecum sepimentum var. arcuata de Folin, 1880: 809 .

Caecum lilianum Hedley, 1903: 603, pl. 29, fig. 7.

Caecum maculata [sic] Habe, 1963: 236, fig. 2.

Caecum berberense Ladd, 1972: 22, 23, pl. 5, figs 11, 12.

Caecum septimentum [sic] Lightfoot, 1992: 1, 4, figs 2, 3.

Caecum gracile Sasaki, 2008: 169, fig. 9E, F.

Caecum sepimentum: Albano & Pizzini 2011: 6–9, fig. 4A–D.

Caecum sepimentum: Pizzini et al. 2013: 5–8, figs 8A–D, F, G, 19K, 20A–D.

Type locality: Mauritius I.

Type material examined: Lectotype (MNHN-IM- 2000 - 24907) selected by Pizzini et al. (2013: 5, fig. 8D).

Material examined: SOUTH AFRICA: Eastern Cape: 1 sh, Mzamba, beachdrift, leg. R.Kilburn & D. Herbert 12–30.v.1986 (NMSA A6285); KwaZulu-Natal: 5 lv and 12 sh, Aliwal Shoal ± - 16 m, hand-dredged sand (NMSA S5943); 5 lv and 39 sh, Landers Reef, off Scottburgh area, - 33 m, SCUBA (JPM); 3 lv and 135 sh, Aliwal Shoal, off Umkomaas area, - 27 m, SCUBA, i.1988 (JPM); 7 sh, Landers Reef, off Scottburgh area, - 45 m, SCUBA, iv.1988 (JPM); 1 sh, Mission Rocks, N of St. Lucia, v.1989 (JPM); 5 lv and 19 sh, Aliwal Shoal, off Scottburgh, - 25–27 m, sand and reef debris (NMSA S7186); 34 lv and 55 sh, Aliwal Shoal, off Scottburgh, - 14 m, underwater pump (NMSA S8627); 6 lv and 1 sh, Aliwal Shoal, Cracker Reef, - 23 m (NMSA E7168); 3 sh, Park Rynie, - 50 m, coarse sand, 2–25.xi.1976, ex C.S.I.R. water Res. 1977 (NMSA B240); 2 lv and 9 sh, Aliwal Shoal, off Scottburgh, - 10–20 m, sand (NMSA S6822); 3 lv and 5 sh (1 juv), Landers Reef (off Park Rynie), - 30 m, slightly muddy sand (NMSA E1671); 1 sh, Landers Reef, - 20–30 m, sorted from stone washings (NMSA E2317); 7 lv and 5 sh, Aliwal Shoal, - 9–15 m (NMSA E6165); 7 lv (1 juv) and 5 sh, Aliwal Shoal, off Scottburgh, - 10 m, sand and reef debris (NMSA S8247); 11 sh, off Sodwana Bay, sediment at base of drop-off in canyon, - 46 m, Marine Geoscience Unit, 7.xii.1992 (NMSA V2626); 8 lv and 4 sh, Kosi Bay, main reef, 1–4 km S of estuary mouth, - 23–24 m, underwater pump (NMSA S2918); 16 lv and 47 sh, Leadsman Shoal, outer portion - 24–26 m, sorted from stone washings (NMSA E2677); 2 lv, Leadsman Shoal, raggie reef, - 8–14 m, a mixed algal and coral reef, 1–2 km N of Leven Point, sorted from stone washings (NMSA E2725); 3 lv and 9 sh, Leadsman Shoal, outer portion, - 25–28 m (NMSA E6929); 2 lv, Leadsman Shoal, - 15 m, scrapings + broken down coral rubble (NMSA E2273); 1 lv and 8 sh, Stn ZB7, off Boteler Point (27°00.5'S 32°54.7'E), - 50 m, coral rubble, dead Lithothamnium, R.V. Meiring Naudé, 6.6.1987 (NMSA E1588); 1 sh, Sodwana Bay, two-mile reef (outer edge), sorted from stone washings, ­ 15–17 m (NMSA D5304); 5 sh, Stn ZA21, SE of Kosi R. mouth (26°55.2'S 32°55.0'E), ­ 50 m, fine sand, dredged NMDP, 8.vi.1987 (NMSA E1516); 1 lv and 16 sh, between Bhanga Neck and Kosi Bay, reef off marker, 13 km North, - 13 m, hand-dredged sand (NMSA S3118); 1 sh, Stn ZM7, off Cape Vidal (29° 07.1S 32° 36.6'E), - 145 m, medium sand, dredged, NMDP, 10.vi.1988 (NMSA E7550); 1 sh, Stn ZF8, off Hully Point (27°.4'S 32°46.4'E), ­ 40 m, fine sand, dredged, NMDP, 5.vi.1987 (NMSA E1658); 4 sh (worn), Stn ZK2, off Gipsy Hill (27°48.8'S 32°38.4'E), - 50 m, dredged, NMDP, 8.vi.1988 (NMSA 4184); 1 sh (worn), Stn ZA19, off Kosi estuary (26°56.7'S 32°54.7'E), - 50 m, coarse sand, shells, dredged, NMDP, 8.vi.1987 (NMSA E1630); 22 sh (worn), Stn ZO4, coarse sand with mud, dredged, NMDP, 12.vi.1988 (NMSA E5197); 1 lv and 2 sh, Sodwana Bay, two-mile reef, sorted from stone washings, - 10–15 m (NMSA E769); 2 lv and 4 sh, Kosi Bay, main reef, 1–4 km S of estuary mouth, - 20 m, underwater pump (NMSA S2526); 3 lv, off Kosi Bay, main reef, 1–2 km S of estuary, sorted from stone washings, - 9–17 m (NMSA D9833); 1 sh, off Kosi Bay, main reef, 1–4 km S of estuary, - 9–17 m (NMSA D9267); 1 sh, Stn ZN7, S.E. of Mission Rocks (28°17.5'S 32°34.2'E), - 100 m, medium sand, dredged, NMDP, 11vi.1988 (NMSA E5191); 2 sh (worn), Stn ZK4, NE of Gipsy Hill (27°45.2'S 32°39.8'E), - 110 m, sponge, stones, dredged, NMDP, 8.vi.1988 (NMSA E5204); 14 sh, Stn ZB 5, off Boteler Point (27°00.0'S 32°55.2'E), - 70 m, coral rubble, dredged, NMDP, 6.vi.1987 (NMSA E1566); 2 lv and 21 sh, Stn ZN1, SE of Mission Rocks (28°17.5'S 32°32.6'E), - 50 m, old coral rubble, Lithothamnium, dredged, NMDP, 10.vi.1988 (NMSA E6252); 2 sh, Stn ZK1, off Gipsy Hill (27°49.7'S 32°38.2'E), ­ 47–50 m, fine sand, dredged, NMDP, 8.vi.1988 (NMSA E5663); 2 lv and 1 sh, between Bhanga Neck and Kosi Bay, reef off marker 13 North, - 8 m, underwater pump (NMSA S2738); 2 lv and 1 sh, Leadsman Shoal, main portion of coral reef, - 7–11 m (NMSA E6774); 12 lv and 11 sh, Kosi Bay, main reef, 1–4 km S of estuary mouth, - 20–22 m, underwater pump (NMSA S2010); 14 sh, Stn ZE2 off Rock I. (27°17.2'S 32°48.0'E), - 62 m, sandstone, coral, marine growths, dredged, NMDP, 5.vi.1987 (NMSA E1542); 1 sh, Stn ZN1, SE of Mission Rocks (28°17.5'S 32°32.6'E), - 50 m, old coral rubble, Lithothamnium, dredged, NMDP, 10.vi.1988 (NMSA E7493); 46 sh, Stn ZK3, off Gipsy Hill (27°47.4'S 32°38.9'E), - 65–70 m, broken shell, dredged, NMDP, 8.vi.1988 (NMSA E5893); 12 lv and 19 sh, Kosi Bay, main reef, 1–4 km S of estuary mouth, - 18 m underwater pump (NMSA S4252); 2 sh, Stn ZK6, off Gipsy Hill (27°48.9'S 32°38.4'E), ­ 50 m, fine sand, dredged, NMDP, 8.vi.1988 (NMSA E7497); 7 sh, Stn Zl5, NE of Leven Point (27°55.0'S 32°38.8'E), - 250 m, coarse sand, dredged, NMDP, 9.vi.1988 (NMSA E5883); 1 sh, Stn ZM9, off Cape Vidal (28°08.3'S 32°36.9'E), - 200 m, sponge rubble, dredged, NMDP, 11.vi.1988 (NMSA E5887); 1 sh, off Kosi Bay, main reef 1–2 km S of estuary, sorted from stone washings, - 9–17 m (NMSA D9892); 1 sh, Leadsman Shoal, Raggie Reef, - 8–12 m, mixed algal and coral reef, 1–2 km N of Leven Point (NMSA E6814); 4 sh, off Richards bay, ­ 50 m, fine quartzite, broken sh, ex C.S.I.R. Water Research (NMSA A6112); 1 sh, off Sodwana Bay, - 50 m, shell grit, ex C.S.I.R. Water Research (NMSA A5087); 7 sh, off Kosi Bay, - 50 m, shell grit, ex C.S.I.R., 1975 Water Research (NMSA A4963); 1 sh, off Durban Bluff, 10 km SE of Cooper Lighthouse, - 210 m, cone dredge, ex C.S.I.R., 1974 (Ecor 5) (NMSA A2993); 26 sh, off Kosi Bay, - 100 m, ex C.S.I.R. Water Research 1977 (NMSA A5898); 20 sh, off Sodwana Bay, - 100 m, ex C.S.I.R., 1976, (NMSA A5553); 3 sh, off Kosi Bay, - 50 m, shell grit, ex C.S.I.R. Water Research (NMSA A5964).

Doubtful: SOUTH AFRICA: Eastern Cape: 1 sh (juv), Mzamba, beachdrift, leg. R. Kilburn & D. Herbert 12–30.v.1986 (NMSA W9680) ; 3 sh, Mzamba, beachdrift, leg. R. Kilburn & D. Herbert 12–30.v.1986 (NMSA D3021) ; 1 sh, Mtentu R. mouth, iv.1982 (JPM 2414) ; KwaZulu-Natal: 1 sh (larval), Landers Reef, off Scottburgh Area, - 45 m, SCUBA, iv.1988 (JPM) ; 2 sh (worn), T. O. Strand, 2 km N of Port Edward (NMSA E1664) .

Additional material examined: RÉUNION: 10 lv and 3 sh, Cap La Houssaye, dived, - 8–17 m, varied bottom (NMSA K5444) ; 46 lv and 25 sh, Cap La Houssaye, - 6–12 m (NMSA) ; 2 sh, Cap de La Réunion (MP) ; 2 sh, Réunion Is. (MP); TANZANIA: Zanzibar: 33 sh, Uroa (MP); KENYA: 3 sh, Kilifi, lagoon inshore of coral reef, sand from base of coral outcrops, - 4 m (NMSA K7997); SUDAN: Red Sea: 57 sh, Sanghaneb Reef, leg. M. Oliverio 6.ii.1998 (MP) ; 6 sh, leg. M. Oliverio 4.ii.1992 (MP); 23 sh (2 larval), Sha’ab Rumi, leg. M. Oliverio 4.ii.1992 (MP) ; EGYPT: Red Sea: 47 sh, Abu Rhamada, Hurghada, leg. Liverani xi.1991 (MP) ; 2 dd, Sinai, leg. I. Nofroni; JORDAN: 1 sh (juv), Aqaba, leg. Liverani iv.1993; MAURITIUS: 3 sh Pointe Radeau, S of Roches Noires, shell debris, near reef gap (NMSA K9511) ; 2 sh, Riambel Lagoon, beach drift (NMSA K9821) ; 8 sh, Gris Gris, debris on surf beach opposite reef-break (NMSA K9186) ; INDIA: 1 sh, Andaman Is. (MP); JAPAN: 3dd, Amami I. (MP); HAWAII: 5 sh, Pali Coast, Maui I. (MP); 1 lv and 2 sh, SE Mokapu Point, Oahu I. (MP); 1 sh, Lanai I. (MP) ; AUSTRALIA: Queensland: 31 dd, N of Green I., E of Cairns (MP) .

Distribution: Indo­West Pacific; this species is very common on the eastern coast of South Africa (Fig. 15).

Remarks: South African specimens show no relevant differences with respect to the specimens found all over the Indo­West Pacific.